Instructions for use Title SOME REMARKS ON THE

Title
Author(s)
Citation
Issue Date
SOME REMARKS ON THE TAXONOMY OF THE
FUNGUS HYPOCHNUS SASAKII SHIRAI
Matsumoto, Takashi
札幌博物学会会報 = Transactions of the Sapporo Natural
History Society, 13(3): 115-120
1934-06-20
DOI
Doc URL
http://hdl.handle.net/2115/64123
Right
Type
article
Additional
Information
File
Information
Vol.13No.3_005.pdf
Instructions for use
Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP
SOMEREMARKSONTHETAXONOMY
OFTHEFUNGUSHYPOCHNUSSASAKIISHIRAI*
BY
TAKASHI MATSUMOTO
{
松
本
説
)
(
W
i
t
h2 t
e
x
t
f
i
g
u
r
e
s
)
む
の a
sw
e
l
la
sbyt
h
ep
r
e
s
沼n
ta
u
t
h
o
r
<
1
4
'
,
As h
a
sa
l
r
e
a
d
ybeens
t
a
t
e
dbyENDO
somes
p
e
c
i
e
sd
e
s
i
g
n
a
t
e
da
sR
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
iKiiHNi
nP
h
i
l
i
p
p
i
n
e
sand I
n
d
i
aa
r
e
a
s
a
k
i
i
q
u
i
t
ei
d
e
n
t
i
c
a
lw
i
t
ht
h
ef
u
n
g
iwhicha
r
eg
e
n
e
r
a
l
l
yr
e
f
e
r
r
e
dt
oa
sHypochnusS
S
H
I
R
A
I
. Neverth
怠l
e
s
s
,o
b
v
i
o
u
s
l
ymuch s
t
i
l
lr
e
m
a
i
n
st
obedone r
e
g
a
r
d
i
n
gt
h
e
モs
e
n
tp
a
p
e
r Iam u
n
a
b
l
e
taxonomicr
e
l
a
t
i
o
n
s
h
i
po
ft
h
e
s
etwos
p
e
c
i
e
s
. I
nt
h
ep
r
t
od
i
s
c
u
s
sf
u
l
l
ya
b
o
u
tt
h
em
a
t
t
e
r ona
c
怠o
u
n
to
fal
i
m
i
t
e
dnumbero
fp
a
g
e
s
,but
w
i
l
lp
e
r
m
i
t myselfo
n
l
yt
omake ab
r
i
e
fr
e
v
i
e
wo
ft
h
etaxonomyo
ft
h
ef
u
n
g
i
under c
o
n
s
i
d
e
r
a
t
i
o
n
.
I
n myp
r
e
v
i
o
u
sp
a
p
e
r
si
th
a
sbeenc
o
n
f
i
r
m
e
dt
h
a
to
u
rf
u
n
g
i can be d
i
s
t
i
n
c
t
l
ys
e
p
a
r
a
t
e
d fromt
h
ec
a
u
s
a
lf
u
n
g
u
so
f“
Pockenkranl
王h
e
i
t
”ofp
o
t
a
t
ot
u
b
e
r
i
nc
u
l
t
u
r
a
lc
h
a
r
a
c
t
e
r
s
,h
y
p
h
a
lf
u
s
i
o
n
,andt
e
m
p
e
r
a
t
u問 r
e
l
a
t
i
o
n
. Thel
a
t
t
e
rf
u
n
g
u
s
wasi
s
o
l
a
t
e
dfromt
h
es
u
p
e
r
f
i
c
i
a
ls
c
l
e
r
o
t
i
a
lmassofp
o
tばob
l
a
c
ks
p
e
c
k(
o
rb
l
a
c
k
e
r
l
i
n
.
s
c
u
r
f
)f
u
n
g
u
sc
o
l
l
e
c
t
e
dbyt
h
ea
u
t
h
o
ri
n Dahle叶 B
The f
i
r
s
tq
u
e
s
t
i
o
nn
a
t
u
r
a
l
l
ya
r
i
s
e
s whethero
rn
o
tt
h
i
sp
o
t
a
t
oo
r
g
a
n
i
s
m
,
r
e
dt
ot
h
e
i
.e
. No. 1
9a
c
c
o
r
d
i
n
gt
o.
o
u
rn
u
m
e
r
a
t
i
o
n
, canbe u
n
m
i
s
t
a
k
a
b
l
y児島r
t
y
p
es
p
e
c
i
e
sR
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
id
e
s
c
r
i
b
e
dbyKuHN
仁川, i
nasmucha
st
h
eb
i
n
o
m
i
a
l
R
h
i
z
o
c
t
o
n
i
aS
o
!
a
n
ihasbeenusedf
o
ra
l
lt
h
ef
u
n
g
im
o
r
p
h
o
l
o
g
i
c
a
l
l
y moreo
rl
e
s
s
r
e
s
e
m
b
l
i
n
gt
h
eKftHN
’
sp
o
t
a
t
op
a
t
h
o
g
e
n
. Accordingt
oD
u
G
G
A
R
<
5
>whoc
o
n
f
e
n
:
e
d
w
i
t
hP
r
o
f
.K
f
t
H
N
,t
h
en
o
m
e
n
c
l
a
t
o
ro
fR
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
i
,r
e
g
a
r
d
i
n
gt
h
i
sd
i
s
e
a
s
e
i
nt
h
ew
i
n
t
e
ro
f1
8
9
9
1
9
0
0
,i
t wasn
o
t
e
dt
h
a
twhen KftHNf
i
r
s
td
e
s
c
r
i
b
e
dt
h
e
(SchorfoderGrind,"
d
i
s
e
a
s
eοfp
o
t
a
t
oi
nGermanyhel
a
i
de
m
p
h
a
s
i
s
.uponas
c
a
b“
Pod
ぽe
n
l
l
a
t
e
r termed “
throughοutt
h
ew
o
r
l
d
. I
nview o
ft
h
ef
a
c
t
,t
h
e
r
e
f
o
r
e
,i
tseems l
i
k
e
l
yt
h
a
to
u
r
p
o
t
a
t
oorganismmaybei
d
e
n
t
i
c
a
lw
i
t
ht
h
es
p
e
c
i
e
sd
e
s
c
r
i
b
e
dbyK
f
i
H
N
. I
nt
h
i
s
c
o
n
n
e
c
t
i
o
n
,i
ti
sa
l
s
ot
on
o
t
et
h
a
to
u
rp
o
t
a
t
oo
r
g
a
n
i
s
mi
sv
e
r
yc
l
o
s
e
l
yr
e
l
a
t
e
dt
o
t
h
ec
u
l
t
u
r
ed
e
s
i
g
n
a
t
e
da
s P4 i
nmyp
r
e
v
i
o
u
sp
a
p
e
r
n
3
>p
u
b
l
i
s
h
e
dw
h
i
l
e Iwasi
n
S
t
.L
ο
u
i
s
, U. S
.A
.
,t
h
el
a
t
t
e
rb
e
i
n
gi
s
o
l
a
t
e
dfroms
c
l
e
r
o
t
i
ao
fb
l
a
c
ks
p
e
c
ko
f
問 do
nt
h
emarketo
fS
t
. 1心 u
i
s
,1
9
1
8
.
p
o
t
a
t
ot
u
b
e
rs
e
c
u
骨 F
r
o
mt
h
eP
h
y
t
o
p
a
t
h
o
l
句i
c
a
lL
a
b
o
r
a
t
o
r
y
,T
a
i
h
o
k
uI
m
p
e
r
i
a
lU
n
i
v
e
r
s
i
t
y
,F
o
r
m
o
s
a
,J
a
p
a
n
.
i
o
c
.
,V
o
l
.X
I
I
I
,P
t
.3
.1
9
3
4
]
[
T
r
a
n
s
.S
a
p
p
o
r
oN
a
t
.日 胤 t
116
TRANSACTIONS OF THE SAPPORO NATURAL HISTORY So:IETY
意
ト
F
i
g
.1
.A
,Rhizoctonz
・
a .~·o!ani, h
y
p
h
a
ea
n
ds
c
l
e
r
o
t
i
a
lc
e
l
l
s
;B,o
u
rf
u
n
g
u
s
,
h
y
p
h
a
ea
n
ds
c
l
e
r
o
t
i
a
lc
e
l
l
s
.(
R
e
d
u
c
e
dt
o~ f
r
o
mc
a
m
e
ほ l
u
c
i
da
d
r
a
w
i
n
g
s
,A
p
o
c
h
.Z
e
.
i
s
s
. 7x4ロ
・
)
Thev
e
g
e
t
a
t
i
v
ec
h
:
i
r
a
c
t
e
r
so
ft
h
ef
u
n
g
u
s正f
y
p
o
c
h
n
u
sS
a
s
a
k
i
ia
r
ea
l
m
o
s
ts
i
m
i
l
a
r
t
ot
h
o
s
eo
fR
h
i
z
o
c
t
o
n
i
a
si
nmanyr
e
s
p
e
c
t
s
. Themyceliumo
ft
h
ef
o
r
m
e
ri
sc
h
:
i
r
a
c
t
e
r
i
z
e
dbyac
o
n
s
t
r
i
c
t
i
o
n wherebranchingo
c
c
u
r
s
, and a septumn
e
a
rt
h
econs
t
r
i
c
t
i
o
n
, becoming d
e
e
p
l
yc
o
l
o
r
e
dandbranchinga
p
p
r
o
x
i
m
'
.
l
t
e
l
ya
tr
i
g
h
ta
n
g
l
e
s
t
ot
h
emain hypha whenm
a
t
u
r
e
,w
h
i
l
ei
nt
h
eyoungers
t
a
g
ei
ti
sc
o
l
o
r
l
e
s
sand
)
. The s
c
l
e
r
o
t
i
aa陀 ge配 r
a
l
l
y more
i
n
c
l
i
n
e
di
nt
h
ed
i
r
e
c
t
i
o
no
fgrowth (
f
i
g
.1
o
rl
e
s
sf
l
a
t
t
e
n
e
d
,i
r
r
e
g
u
l
a
r
, brown, and f
a
i
r
l
yh
o
m
o
g
e
n
e
o
:
.
i
s and p
:
i
r
a
p
l
e
c
t
e
n
chymatousi
ns
t
r
u
c
t
u
r
e
, composing o
ft
h
ec
e
l
l
sc
o
n
c
o
l
o
r
o
c
r
s with t
h
es
u
r
f
a
c
e
.
巴t
ョt
i
v
es
t
a
g
e
,t
h
ef
u
n
5
u
s HyAlthough manys
i
m
i
l
a
r
i
t
i
e
se
x
i
s
ti
nt
h
eveg
sd
i
s
t
i
n
c
t
l
yd
i佐 r
e
n
tfromR
f
t
i
z
o
c
t
o
m
・
d S
o
l
a
n
ii
n somemorphopochnus S
a
s
a
k
i
ii
l
o
g
i
c
a
l and p
h
y
s
i
o
l
o
g
i
c
a
lc
h
a
r
a
c
t
e
r
s
,s
of
a
ra
sac
o
m
p
:
i
r
i
s
o
ni
smade between
rr
i
c
es
t
r
a
i
n(
N
o
.1
)andthepotato Rhizoctonia(No.1
9
)
. Forc
:
:
i町 e
n
i
e
n
c
e
,
oコ
some o
ft
h
e morec
o
r
:
t
r
a
s
t
i
n
gf
e
a
t
u
r
e
sa
s usmlly found a
r
eshowni
nt
h
ef
o
l
lowing t
a
b
l
e:
Hypochnus Sasakii
A. M
o
r
p
h
o
l
o
g
i
c
a
lc
h
a
r
a
c
t
e
r
s(
f
i
g
.1
)
I. H
yphae:
5
1I μ
,mostly6-8μ
.
2. S
c
!
e
r
o
t
i
a
lc
e
l
l
s(
n
e
a
rt
f
t
e炉 r
i
少h
e
r
y
)
:
7-18x22-37μ
.
R
h
i
z
o
<
t
.
.
o
n
i
aSolani
A. Morphological c
h
:
i
r
a
:
:
:
t
e
r
s
I. H
yphae:
7-13μ
, mostly 8
-12p
.
.
2. S
d
e
r
o
t
i
a
lc
e
l
l
s(
d
o
.
)
:
15-26×26-50μ
.
MATSUMOTO: O N THE TAXONOMY OF め'
f
o
e
h
n
t
t
sS
a
s
a
k
i
i
1
1
7
B. P
h
y
s
i
o
l
o
g
i
c
a
lc
h
a
r
a
c
t
e
r
s
B. P
h
y
s
i
o
l
o
g
i
c
a
lc
h
a
r
a
c
t
e
r
s
On会h
eη:
o
lγe
d
C
z
α
r
p
e
k
'
sα
g
α
γ:
Mostoft
h
es
t
r
a
i
n
se
x
h
i
b
i
t“
Peach
r“
S
c
a
r
l
e
t” i
n 7d
a
y
s
,
r
e
d” o
showingi
n
c
r
e
a
s
e
da
l
k
a
l
i
n
i
t
y
.
2. On b
r
o
m
c
r
e
s
o
l抑 r
p
l
e
D
u
g
g
a
〆s
α
z
α
γ:
A
f
t
e
r 8 days a
l
lt
h
ec
u
l
t
u
r
e
sb
e
come “
Mustardyellow" on a
c
counto
ft
h
es
h
i
f
t
i
n
gt
oa
c
i
ds
i
d
e
.
I.
I.
3・ Onαη£
;
£
!机 eb
l
u
e
-C
z
a
p
e
k
'
sα
i
g
a
r
:
A moderately s
t
r
o
n
g growth i
n
a
l
lt
h
ec
u
l
t
u
r
e
s
.
4
. Ont
a
n
n
£
cαc
i
d
p
o
t
α
,t
oα
g
αr
:
More o
rl
e
s
st
o
l
e
r
a
n
tt
ot
h
ea
c
i
d
.
5
. F
u
s
i
o
no
fh
y
p
l
i
a
e
:
Fusion t
a
k
e
sp
l
a
c
e between any
s
t
1
τ百
L
i
n
so
fH
ypochnus S
α
s
αk
i
i
.
6
. T
e
m
p
e
r
a
t
u
r
er
e
l
a
t
i
o
n:
Thesea
r
er
a
p
i
d
g
r
o
w
i
n
gf
u
n
g
i
,g
i
v
i
n
g most l
u
x
u
r
i
a
n
t growth i
n2
days a
t 28-3I。
c
.
Onp
h
e
n
o
lr
e
d
C
》a
p
e
k
'
sa
g
a
r
:
T
h
i
sc
u
l
t
u
r
ebecomes“
B
i
t
t
e
r
s
w
e
e
t
orange
,
” thusshowingl
e
s
sa
l
k
a
l
i
n
e thant
h
ef
o
r
m
e
r
.
On b
r
o
m
c
r
e
s
o
l pu
ヴ'l
e
D
z
必'
l
{
t
r
r
'
s
a
g
a
r
:
This c
u
l
t
u
r
e becomes “
Antimony
yellow
,
” thus showing s
l
i
g
h
l
y
l
e
s
sa
c
i
dt
h
a
nt
h
ef
o
r
m
e
r
.
3
. Ona
n
i
l
i
n
eb
l
u
e
C
》a
p
e
k
’
sa
g
a
r
:
T
h
i
sf
u
n
g
u
si
sv
e
r
ys
e
n
s
i
t
i
v
et
o
t
h
ed
y
e
.
4
. Ont
a
m
z
i
ca
c
i
d
-C》a
p
e
k
'
sagar:
Verys
e
n
s
i
t
i
v
et
ot
h
ea
c
i
d
.
5
. 乃t
s
i
o
ne
fめア加に
Nof
u
s
i
o
nt
a
k
e
sp
l
a
c
ei
nc
r
o
s
st
e
s
t
s
w
i
t
h any o
ft
h
ef
o
r
m
e
r
.
6
. T
t
T
h
i
s i
s a slow-growing f
u
n
g
u
s,
showingnomarkedd
i
白e
r
e
n
c
ei
n
tt
e
r
n
growthevena
f
t
e
r2 daysa
p
e
r
a
t
u
r
e
s223I0C
.Theoptimum
i
slowert
h
a
nt
h
a
to
ft
h
ef
o
r
m
e
r
.
2.
ρmvf
I
nadditiont
ot
h
e
s
ef
a
c
t
s
,i
ti
sa
l
s
o
t
on
o
t
et
h
a
td
i
r
e
c
ta
t
t
a
c
k ofl
e
a
v
e
sby
R
l
z
i
z
o
c
t
o
n
i
aS
o
l
a
n
ii
sr
a
t
h
e
ri
n
f
r
e
q
u
e
n
t
witht
h
ee
x
c
e
p
t
i
o
n of such p
l
a
n
t
sa
s
l
e
t
t
u
c
e
,c
a
b
b
a
g
e
,e
t
c
.
, wherea? i
n our
c
a
s
el
e
a
v
e
sa
r
emoref
r
e
q
u
e
n
t
l
ya
t
t
a
c
k
e
d
by t
h
ef
u
n
g
u
s
.
A comparison o
ft
h
e two s
p
e
c
i
e
s
i
nt
h
ep
e
r
f
e
c
ts
t
a
g
ea
l
s
or
e
v
e
a
l
s some
d
i
f
f
e
r
e
n
c
e
s
,p
r
o
v
i
d
e
dC
o
r
t
i
c
i
u
mvagum
B
.& C
.d
e
s
c
r
i
b
e
d by BURT<~> i
st
h
e
F
i
g
.2
.
C
o
r
t
i
e
加'
!
lv
agumか
t
p
p
e
r
)
,× 800.
a
,f
r
o
ms
p
e
c
i
m
e
no
np
o
t
a
t
o;c
,f
r
o
m
s
p
e
c
i
m
e
no
ne
a
r
t
h;d
,f
r
o
ms
p
e
c
i
m
e
no
nw
o
o
d
. (
R
e
p
r
o
d
u
c
e
df
r
o
m
B
u
R
T
l
4
>
)O
u
rf
u
n
g
u
s(
l
o
w
e
r
)
,x500.
(
S
. HIRANE d
e
!
.
)
け
~
T
R
A
N
S
A
C
T
I
O
N
SO
FT
H
ES
A
P
P
O
R
ONATURALH
I
S
T
O
R
YS
o
c
J
E
τ
y
I
I
8
p
e
r
f
e
c
ts
t
a
g
eo
fR
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
i
. Fori
n
s
t
a
n
c
e
,t
h
e number o
fb
a
s
i
d
i
o
s
p
o
r
e
s
o
f r
t
i
c
i
・
umvagum v
a
r
i
e
s from 4 t
o6
, thoughi
nt
h
ei
l
l
u
s
t
r
a
t
i
o
n of RoLFs
(
r
e
p
r
o
d
u
c
e
d by Gussm
が
の
) t
h
eb
a
s
i
d
i
aa
r
e2t
o4 s
t
e
r
i
g
r
n
a
t
e
d
, wl
町 e
a
si
n our
c
a
s
et
h
eb
a
s
i
d
i
o
s
p
o
r
e
s vary from 2 t
o 4 on each b
a
s
i
d
i
u
r
n
, and a
r
es
l
i
g
h
t
l
y
s
h
o
r
t
e
r but broader t
h
a
nt
h
e former (
f
i
g
.2
)
. The c
h
a
r
a
c
t
e
r
i
s
t
i
cf
e
a
t
u
r
e
sa
s
d
e
s
c
r
i
b
e
d by s
e
v
e
r
a
li
n
v
e
s
t
i
g
a
t
o
r
sa
r
e shown i
nt
h
ef
o
l
l
o
w
i
n
gt
a
b
l
e:
ο
Hyp
配 h
m.
臼
rgum
C
o
r
t
z
c
i
u
m℃α
S
αs
a
k
i
i
S
A
W
A
D
A
(
l
S
)
IIKATA& 酌OM
Thea
u
t
h
o
r
C
U
)
B
a
s
i
d
i
a
:
J
o
1
0
2
0X7
.
5
II(
1
0
1
5x7
9(
J
o
5
.
6
1
2x5
8(
J
o
1
0
1
6x
8
9μ
S
t
e
r
i
g
m
a
:
6
1
6
μ
,
4
6s
t
e
r
i
g
m
a
t
e
d
4
.
5
7x2
3μ
.
2
4s
t
e
n
g
m
a
t
e
d
2
・
5
8
.
4x0
.
8
2
.
8μ
5
8X
2
;
2
2・
7μ
,
2
4s
t
e
r
i
g
m
at
e
d
E
a
s
i
d
i
o
s
p
o
r
e
s:
8
1
4x4
6μ
8
・E1
x5・6
.
5件
6
.
5
8
.
4x4
.
8
8
.
4μ
6
r
ox4
7件
!仰
I
ti
sa
l
s
ot
on
o
t
et
h
a
tw
h
i
l
e Iwasworking with R
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
ii
nS
t
.
,o
r
i
g
i
n
a
t
e
d from C
o
r
t
i
c
i
u
r
ns
t
a
g
e on l
i
r
n
a
Louis Ifound t
h
a
t as
t
r
a
i
n
,i
.
e
. B3*
beans grewb
e
t
t
e
ra
t lowert
e
m
p
e
r
a
t
u
r
e
s(
c
a
.24°
C
.)
. Thistemperaturer
e
l
a
t
i
o
n
i
sa
p
p
a
r
e
n
t
l
yd
i任e
r
e
n
tfrom t
h
a
to
fourf
u
n
g
i and r
a
t
h
e
rr
e
f
e
r
a
b
l
et
ot
h
a
to
fO~Ir
No. 1
9s
t
r
a
i
n(
R
.
んi
z
o
c
t
o
n
i
aS
o
l
a
n
i
)
.
Although t
h
em
o
r
p
h
o
l
o
g
i
c
a
ld
i民 r
e
n
c
e
s between t
h
e
s
e two f
u
n
g
ii
nt
h
e
p
e
r
f
e
c
ts
t
a
g
ea
r
en
o
ts
u日c
i
e
n
tenought
ow
a
r
r
a
n
t ac
l
a
s
s
i
f
i
c
a
t
i
o
no
four fung~1s
a
sad
i
s
t
i
n
c
ts
p
e
c
i
e
s
,i
njudging from t
h
ev
e
g
e
t
a
t
i
v
ec
h
a
r
a
c
t
e
r
so
f ourf
u
n
g
:
.
i
s
o
r
t
£
c
i
u
mvagum, and shouldbes
e
p
a
r
a
t
e
d
t
h
i
s cann
o
tbei
n
c
l
u
d
e
di
nt
h
es
p
e
c
i
e
sC
from t
h
el
a
t
t
e
ra
sa d
i
s
t
i
n
c
ts
p
e
c
i
e
sr
a
t
h
e
rt
h
a
na
s ab
i
o
l
o
g
i
cf
o
r
m
.
I
nt
h
i
sc
o
n
n
e
c
t
i
o
ni
t may be n
e
c
e
s
s
a
r
yt
o make a b
r
i
e
fr
e
v
i
e
w regardin~
t
h
es
p
e
c
i
e
s砂 i
p
o
c
h
n
u
sS
.
ぬ£
n (C
o
r
t
i
c
i
u
mS
o
l
a
n
i
)
,s
i
n
c
et
h
i
sb
i
n
o
m
i
a
l has been
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
i by s
e
v
e
r
a
li
n
v
e
s
t
i
g
a
t
o
r
s
,p
a
r
t
i
c
u
l
a
r
l
y by European
used f
o
rR
w
o
r
k
e
r
8
l
'1
1
>
. The s
p
e
c
i
e
s concerned was f
i
r
s
td
e
s
c
r
i
b
e
d by PRILLIEUX and
DELACROIXi
n1
8
9
1
,abouttwodeca仁l
e
sa
f
t
e
rC
o
r
t
i
c
i
・
umvagumhadbeen r
e
p
o
r
t
e
d
,
a
s ac
a
u
s
a
l fungus onp
o
t
a
t
os
t
e
m
, although no c
o
n
n
e
c
t
i
o
nwass
u
s
p
e
c
t
e
dwith
Rhi
・
z
o
c
t
o
n
i
aS
o
l
a
n
i
, already known t
o occuri
n Europe. The d
i
a
g
n
o
s
i
s oft
h
e
ョr
i
s
o
ncould h
a
r
d
l
y be made
fungus was s
oi
m
p
e
r
f
e
c
t
l
yd
e
s
c
r
i
b
e
d
,t
h
a
t a comp
s
a
t
i
s
f
a
c
t
o
r
i
l
y except t
h
a
tt
h
eb
a
s
i
d
i
o
s
p
o
r
e
s appeart
o agree more c
l
o
s
e
l
y with
t
h
o
s
eo
fo
u
r
s
,a
sf
a
ra
s can bejudged from t
h
emeasurement oft
h
es
p
o
r
e
s
c
1
7
'
.
*T
h
i
sm
a
t
e
r
i
a
lw
a
se
x
a
m
i
n
e
db
yD
r
.BURTa
n
dd
e
t
e
r
m
i
n
e
dt
ob
ei
d
e
n
t
i
c
a
lw
i
t
hCorticiumv
a
,
1
,
r
i
:
m
.
(
c
f
.t
h
ep
a
p
e
r
(
1
3
)
入
M
A
T
S
U
M
O
T
O
: ONT
H
ET
A
X
O
N
O
M
YO
F巧pochnusSasaldi
I1
9
Asam
a
t
t
e
ro
ff
a
c
t
,h
o
w
e
v
e
r
,i
ti
st
on
o
t
et
h
a
tt
h
es
p
e
c
i
e
sC
o
r
t
i
c
i
m
nS
o
l
a
n
i
e
s
c
r
i
b
e
dbyB
R
0
0
K
s
<
2
>h
a
sb
a
s
i
d
i
o
s
p
o
r
e
smeasuring8・14x
(
H
y
p
o
c
!
t
n
u
sS
o
!
a
n
i
)d
e
r
f
e
c
t
l
ya
g
r
e
e
sw
i
t
ht
h
a
to
ft
h
eBURT
’
sC
o
r
t
i
c
i
m
nvagum. More
4-6μ whichp
r
e
c
e
n
t
l
y
,i
nr
e
g
a
r
dt
ot
h
en
o
m
e
n
c
l
a
t
u
r
e o
ft
h
ep
e
r
f
e
c
t form o
ft
h
ep
o
t
a
t
o
R
h
i
z
o
c
t
o
n
i
aB
R
A
U
N
(
t
>n
o
t
i
c
e
st
h
a
ti
ti
sn
o
ty
e
tq
u
i
t
ec
e
r
t
a
i
nwhethert
h
ef
u
n
g
u
s
s
h
o
u
l
d ber
e
g
a
r
d
e
da
saHypochnuso
raC
o
r
t
i
c
i
u
m
,o
rwhethert
h
es
p
e
c
i
f
i
cname
s
h
o
u
l
d bevagumo
rS
o
l
a
n
i
. Howevert
h
a
tmayb
e
,i
nviewo
ft
h
eo
p
i
n
i
o
n
s
g
i
v
e
nbymanyi
n
v
e
s
t
i
g
a
t
o
r
supt
ot
h
ep
r
e
s
e
n
tt
i
m
e
,i
tseemsl
i
k
e
l
yt
h
a
t再y
p
o
c
h
n
u
s
sc
o
n
s
i
d
e
r
e
dt
obe i
d
e
n
t
i
c
a
lw
i
t
hC
o
r
t
i
c
i
u
mvagum.
S
o
!
a
n
ii
Thent
h
enexta
t
t
e
n
t
i
o
ni
sd
i
r
e
c
t
e
dt
ot
h
eq
u
e
s
t
i
o
nwhether o
rn
o
tt
h
e
r
e
n
a
r
eanyo
t
h
e
rs
p
e
c
i
e
sb
e
a
r
i
n
g ac
l
o
s
e
rr
e
s
e
m
b
l
a
n
c
et
ot
h
ep
r
e
s
e
n
tf
u
n
g
u
s
. I
viewo
ft
h
el
i
t
e
r
a
t
u
r
ea
v
a
i
l
a
b
l
ea
tp
r
e
s
e
n
ti
twouldseem t
h
a
tourf
u
n
g
u
si
sv
e
r
y
c
l
o
s
e
l
yr
e
l
a
t
e
dt
oC
o
r
t
i
c
i
u
mS
t
e
v
e
n
s
i
iBURT
仏 明 (o
rC
o
r
t
i
c
i
u
mk
o
l
e
r
o
g
a(CooKE)
v
.HδHN. a
c
c
o
r
d
i
n
gt
o WOLFandB
A
c
t
t
(
2
0
>
)
, ac
a
u
s
a
lf
u
n
g
u
so
f hypochnoseo
f
pomaceousp
l
a
n
t
s
,C
i
t
r
u
s
,c
r
o
t
o
n
s
,r
o
s
e
s
,e
t
c
. Thef
u
n
g
u
s οr
t
z
・
ciumS
t
e
v
e
n
s
i
ii
s
c
h
3
.
r
a
c
t
e
r
i
z
e
dbyh
a
v
i
n
gr
o
u
n
d
i
s
ho
roblongs
c
l
e
r
o
t
i
au
s
u
a
l
l
y3o
r4m mi
nd
i
a
m
e
t
e
r
and c
h
e
s
t
n
u
tbrowni
nc
o
l
o
r
, composed o
fcompactlywovenmasses o
fs
w
o
l
l
e
n
i
r
r
e
g
u
l
a
rhyphae whicha
r
e homogeneoust
h
r
o
u
g
h
o
u
tt
h
et
i
s
s
u
e
s
,d
i
r
t
yp
i
n
k
i
s
h
b
u
f
f
,membranaceousf
r
u
c
t
i
f
i
c
a
t
i
o
n
sw
i
t
hs
p
o
r
e
smeasuring 1
0
.
5
1
1
.
6×4・
7
5
.
8μ
(STEVENS & H
A
L
L
(
l
吋 or 8寸 IX 3-4μ (BURT刊
)
, and mycelial rhizomorphic
s
t
r
u
c
t
u
r
e
se
x
t
e
n
d
i
n
gl
e
n
g
t
h
w
i
s
eo
ft
h
et
w
i
g
sandp
e
t
i
o
l
e
s
. Fromt
h
ed
e
s
c
r
i
p
t
i
o
n
s
i
twould seem t
h
a
tt
h
i
sf
u
n
g
u
ssomewhatd
i佐 r
sfrom o
u
r
si
nh
a
v
i
n
gc
h
a
r
a
c
t
e
r
i
s
t
i
cr
h
i
z
o
m
o
r
p
h
i
cs
t
r
u
c
t
u
r
e
s
,i
np
r
o
d
u
c
i
n
gs
c
l
e
r
o
t
i
amostly on t
w
i
g
s
, and i
n
h
o
s
tr
e
l
a
t
i
o
n
. However,botha
r
ev
e
r
yc
l
o
s
e
l
yr
e
l
a
t
e
dt
oeacho
t
h
e
ri
n Rhizoc
t
o
n
i
a
l
i
k
e hyphaln
a
t
u
r
e
,s
c
l
e
r
o
t
i
a
ls
t
r
u
c
t
u
r
e
,f
r
u
c
t
i
f
i
c
a
t
i
o
n
,modeo
fp
e
n
e
t
r
a
t
i
o
n
,
andi
nr
e
l
a
t
i
o
nt
ot
e
m
p
e
r
a
t
u
r
e and h
u
m
i
d
i
t
y
. As r
e
g
a
r
d
st
h
em
a
t
t
e
rs
t
u
d
i
e
s
a
r
es
t
i
l
li
np
r
o
g
r
e
s
s
,s
ot
h
a
tf
u
r
t
h
e
ro
b
s
e
r
v
a
t
i
o
n
sw
i
l
lbep
u
b
l
i
s
h
e
dl
a
t
e
r
. I
n
c
o
n
c
l
u
s
i
o
n I may say t
h
a
ti
t wouldbeb
e
t
t
e
rt
onameo
u
rf
u
n
g
u
sC
o
r
t
i
c
i
u
m
S
a
s
a
k
i
l
.
1
5
>i
n
s
t
e
a
do
fHypochnus S
a
k
a
k
i
iu
n
t
i
lf
u
r
t
h
e
ra
l
t
e
r
a
t
i
o
ni
sn
e
e
d
e
d
.
B
i
b
l
i
o
g
r
a
p
h
y
(
1
)B
R
A
U
NH.: D
e
rW
u
r
z
e
l
t
o
t
e
rd
e
rK
a
r
t
o
偽 IR
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
iK
.M
o
n
o
g
r
.z
u
mP
f
l
a
n
z
e
n
s
c
h
u
t
z
9
3
0
.(
c
ι
R
.
A
.
M
.9:
7
3
9・ 1
9
3
0
)
5
,1
3
6・
P
.・1
(
2
)B
R
O
O
K
S
,E
.
T
.
:P
l
a
n
tdi民 a
s
e
s
.p
p
.2
6
3
2
6
5
・1
9宜8
.
(
3
)B
U
R
T
,E
.A
.
:T
h
eT
h
e
l
e
p
h
o
r
a
c
e
a
eo
fN
o
r
t
hA
m
e
r
i
c
a
.I
.A
n
n
.M
o
.B
o
t
.G
a
r
d
e
nI:1
8
5
2
2
8
.
1
9
1
4
・
{
4
)一
一
ー
一
一
一
・ zTheThelephoraceaeo
fN
o
r
t
hA
m
e
r
i
c
a
.X
V
.C
o
r
t
i
c
i
u
m
.I
b
i
d
.1
3:2
9
5
2
9
8
.
1
9
2
6
.
TRANSACTIONSOFTHESAPPORONATURALHISTORYSOCIETY
120
{
5
) DUGGAR,B
.M.: R
h
i
z
o
c
t
o
n
i
aCrocorum(
P
E
R
S
.
)DC.andR
.S
o
l
a
n
iKUhn(
C
o
r
t
i
c
i
u
mvagumB
.
&C
.
)w
i
t
hn
o
t
e
sono
t
h
e
rs
p
e
c
i
e
s
.I
b
i
d
.2:4
0
3
4
5
8
. 1
9
1
5・
(
6
) E
:
m
o
,S
.
: Ont
h
eS
c
l
e
r
o
t
:
u
md
i
s
e
a
s
eo
fr
i
c
ep
l
a
n
t
si
nt
h
eP
h
i
l
i
p
p
i
n
e
s
. (
A
r
t
i
c
l
ei
nJ
a
p
a
n
e
s
e
)
J
o
u
r
.P
l
a
n
tP
r
o
t
e
c
t
. 14:2
8
3
2
8
8
.1
9
2
7
. Alsoi
nNogyo
・
kenkyu14:ト 3・ 1
9
3
0
.
(
7
) ERIKSSON,J
・
2 DjeP
i
l
z
k
r
a
n
k
h
e
i
t
e
nd
e
rl
a
n
d
w
i
r
t
s
c
h
a
f
t
l
i
c
h
e
nK
u
l
t
u
r
g
e
w
a
c
h
s
e
. IT
e
i
!
.p
p
. 1481
5
0
. 1
9
2
7
.
(
8
) F
o
R
S
T
E
N
E
I
C
H
l
'
I
E
R
,F
.
: D:eJ
u
g
e
n
d
k
r
a
n
k
h
e
i
t
e
nd
e
rBaumwollei
nd
e
rT
i
i
r
k
e
i
. P
h
y
t
o
p
a
t
h
.Z
e
i
t
s
c
h
r
.3:3
6
7
・417
・ 1931.
(
9
) Gussow,H.T.: B
e
i
t目 gz
u
rKenntnisd
e
sK
a
r
t
o
f
f
e
l
G
r
i
n
d
e
s
.C
o
r
t
i
c
i
u
m vagum B
.e
tC
.v
a
r
.
S
o
l
a
n
iBURT. Z
e
i
t
s
c
h
r
.f
.P
f
l
a
n
z
e
n
k
r
. 16:1
3
5
1
3
7
,T
a
f
.V
I
I
I
. 1
9
0
6
.
臥 T
A
,S
.andHrrmu,T.: On t
h
emodeo
fp
r
i
m
a
r
yi
n
f
e
c
t
i
o
nt
h
r
o
u
g
hs
c
l
e
r
o
t
i
a and r
i
e
l
d
(
1
0
) I
o
b
s
e
r
v
a
t
i
o
n
sont
h
eb
a
s
i
d
i
o
s
p
o
r
ef
o
r
m
a
t
i
o
ni
nHypochnusS
a
s
a
k
i
iS
h
i
r
a
io
fr
i
c
ep
l
a
n
t
.
(
A
r
t
i
c
l
ei
nJ
a
p
a
n
e
s
e
) J
o
u
r
.P
l
.P
r
o
t
. 17:1
7
2
8
. 1
9
3
0
.
{
I
I
) KOTILA,J.E.: A s
t
u
d
yo
ft
h
eb
i
o
l
o
g
yo
fanews
p
o
r
e
.
f
o
r
m
i
n
gR
h
i
z
o
c
t
o
n
i
a
,C
o
r
t
i
c
i
u
mp
r
a
t
i
c
o
l
a
.
。
・1
9
2
9・(c
f
.H
i
s
t
o
r
i
c
a
lpp
・1060-1062)
P
h
y
t
o
p
a
t
h
.1
9:1059-1099
2
) KUHN,J
.
: Kra
此h
e
i
t
e
nd
e
rK
u
l
t
l
時・
e
w
a
c
h
s
e
,i
h
r
e Ursachen und i
h
r
eV
e
r
h
i
i
t
u
n
g
.p
p
.2
2
2
'
量2
8
.
1
8
5
8
.
(
1
3
) MATSUMOTO,T
.
: S
t
u
d
i
e
si
nt
h
ep
h
y
s
i
o
l
o
g
yo
ft
h
ef
u
n
g
i
;X
I
I
.P
h
y
s
i
o
l
o
g
i
c
a
ls
p
e
c
i
a
l
i
z
a
t
i
o
ni
n
R
h
i
z
o
c
t
o
n
i
aS
o
l
a
n
iKuHN. Ann.Mo.B
o
t
. Garden8:6
3
9
6
. 1
9
2
1
.
(
1
4
) M
A
τ'S
U
M
O
T
O
,T
.
,y
AMAMOTO,w
.
,andHIRANE,s
.:Physiol句 yandparasitismofthefungige・
n
e
r
a
l
l
yr
e
f
e
r
r
e
dt
oa
sHypochnusS
a
s
a
k
i
iS
h
i
r
a
i
.I
. Di
偽r
e
n
t
i
a
t
i
o
no
ft
h
es
t
r
a
i
n
s by
meanso
fh
y
p
h
a
lf
u
s
i
o
nandc
u
l
t
u
r
ei
nd
i
f
f
e
r
e
n
t
i
a
lm
e
d
i
a
. J
o
u
r
.S
o
c
.T
r
o
p
.A
g
r
. 4:
3
7
0
3
8
8
. 1
9
3
2
.
{
巧
)
一一一一一一.:Physiology and parasitism of thefungi generally referred to as Hypochnus
S
a
s
a
k
i
iS
h
i
r
a
i
.I
I
. T
e
m
i
:
陪r
a
t
u
r
eandh
u
m
i
d
i
t
yr
e
l
a
t
i
o
n
s
.I
b
i
d
. S:332-345
・ 1933・
{
1
6
) Reviewo
fAppliedMycology7:54・ 1
9
2
8
.
3
0
. 1
8
9
5・
(
1
7
) SACCARno,P.A.: S
y
l
l
.F
u
n
g
. 11 :1
“
{
I
S
) SAWADA,K.: Ont
h
e S
h
i
r
a
k
i
n
u
b
y
o
’・ ofcamphor. (ArticleinJapanese) TaiwanAgr.Exp.
S
t
a
t
i
o
nS
p
e
c
i
a
lB
u
l
l
.N
o
.
・・
4 80p
p
.1
9
1
2
.
.G.: Hypochnose o
f pomaceous f
r
u
i
t
s
. Ann. M
y
c
o
l
. 7:49
・
.
5
9
,
・
(
1
9
) S
T
E
V
E
N
S
,F
.L.andHALL,J
1
9
0
9・
t
宮o
)
WOLF,F.A. andBACH,W.J
.
: Thet
h
r
e
a
db
l
i
g
h
td
i
s
e
a
s
e回 開dbyC
o
r
t
i
c
i
u
mk
o
l
e
r
o
g
a(
C
o
o
k
e
)
Hohn.1onC
i
t
r
u
sandpoma
白 o
u
sp
l
a
n
t
s
. P
h
y
t
o
p
a
t
h
.1
7
:6
8
9
7
0
9
,1
9
2
7
・