I ask Dr. Blakemore

I ask Dr. Blakemore “Pheretima koellikeri”
1) Mystery of “Pheretima koellikeri” distributed only in Kyushu
by Kobayashi 1941:
Pheretima koellikeri” – Amynthas vesiculata
the distribution of Amynthas vesiculata
2) Pheretima koellikeri again
Dr. Blakemore's aims on New genus proposal
Re-description of "Pheretima koellikeri" (Manus koellikeri) by Blakemore 2010
Comparison
Pheretima koellikeri, Manus koellikeri, Metaphire vesiculata
3) Metaphire hilgendorfi / Amynthas tokioensis species complex
4) Slide of addition
10 or more specimens of Metaphire ryukyusensis sent from Dr. Hamada to me
Lecture date 20x14 /5x /1x ; Beginning
day :2014/4/30 The description will be corrected in the future.
Metaphire hilgendorfi / Amynthas tokioensis specise complex in Kobayashi (1941)
Metaphire hilgendorfi / Amynthas tokioensis species complex in Kyushu, Sikoku, Chu-goku, Kinki,
Chu-bu and its surrouding
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鮮
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中
シ
ナ
北
シ
ナ
満
州
国
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樺
太
Saghalien
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南
シ
ナ
South Chaina
korea
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台
湾
Taiwan; Formosa
Hokkaido
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Cheju Island
Touhoku
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済
州
島
North east part of Chaina (Manchuria)
北
海
道
North Chaina
東
北
Mid Chaina
関
東
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中
部
Kantou
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近
畿
Chu-bu
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中
国
Kinki
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四
国
Chu-goku
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隠
岐
Shikoku
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九
州
Oki (Island)
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対
馬
Kyushu
Ph. agrestis (Goto et Hatai)
Ph. communissima (G.et.H.)
Ph. glandularis (G. et. H.)
Ph. gomejimensis Ohfuchi
Ph. hilgendorfi (Mihaelsen)
Ph. irregularis (Goto et Hatai)
Ph. parvicystis (G.et.H.)
Ph . soulensis Kobayashi
Ph . tappensis (Ohfuchi)
Ph. vittata (Goto et Hatai)
Ph. yunoshimensis Hatai
Ph. yamadai Hatai
壱
岐
Tusima (Island)
Disappearing Ph. koellikeri
五
島
Iki (Island)
Amamioshima (Island)
甑
島
Gotou Fukue (Island)
Okinawa (Island)
屋
久
島
Kosikijima (Island)
大
島
Yakusima (Island)
沖
縄
Blakemore (03-11)
Pheretima (Parapheretima) koellikeri Michaelsen, 1928 is herein considered
synonymous with the prior Metaphire vesiculata (Goto & Hatai, 1899), thereby
removing the genus Pheretima sensu stricto from the Japanese list.
Pheretima koellikeri again
dr=fragliche acanthodriline
Prostata→Secretory diverticulum duct
[fragliche→questionable]
Pr=Pheretima-Prostata→Prostatic duct
Pb=Penial borste →Penial setae
Figure above was quoted from Record of the Australian
Museum (2000) Vol. 52: 187-222 Native Earthworms
(Oligochaeta) from Southeastern Australia, with the
Description of fifteen New Species by R.J. Blakemore
as Stephenson (1930) remarked, normal glandular prostates with the usual relations are also present, thus duplicated or
transitional functionality is unlikely. (Blakemore 2013)
I might have misunderstood the penial setae
.
I might have misunderstood the penial setae.
Ventral side (6/7/8) of Metaphire vesiculata
boxed sketch: lhs
spermathecae, 18
rhs prostate and
manicate caecum of
the immature
second type
specimen
6/
7/
8
17/18/19
6/ 7 / 8 /9
16/17/18/19 /20
Metaphire vesiculata
24/25 /26/27
Caeca
Spermatheca
Diverticulum
Pheretima koellikeri
Manus koellikeri
Oligochaeta. Das Tierreich,
10. xxix
Michaelsen, W. 1900a.
A New Earthworm Genus
(Oligochaeta, Megadrili,
Megascolecidae) from Japan
Robert J. Blakemore1
Metaphire vesiculata
Several taxa belonging to
Metaphire hilgendorfi/Amynthas
tokioensis species complex having
full sexual organ in Fukuoka
Kyushu Japan Y. Sugi
5 oder 6 schlanke fingerformige, ///
Manicate
(Manicate)
2 pair in 6/7/8
2 pair in 6/7/8
Michaelsen shows one with
in contrast, the current lectotype
zig-zagging diverticular bulb; had flat, rather compact bulbs on
short only slightly wavy stalks
An attached gland is not described.
Other features　1
Manicate
2 pair in 6/7/8
Œ
iverticular bulb is straight.
There is an attached gland of a
long handle near the base of
each spermathea.
Genital markings absent except for pads around spermathecal pores
pads around spermathecal pores
was signs.
Spermatheca surrounding
Prostatic duct
on 18
on 18
Present (shown in Fig. 1)
Penial seta (Penial　borste)
An attached gland is not described.
attached glands
There are two attached glands
with a long handle near the base
of prostatic duct.
I made an effort to understand the paper of manus by Dr. Blalemore. However, I am
uncertain whether to have understood the paper enough.
of "Manus"
one a posteriorFurther
amputeereading
mature, solving
previously
dissected with most of
internal organs removed such as gizzard, one a posterior amputee
mature,
内臓の大部分がgizzard both lhs spermathecae, and primary male
organs except for part of 18lhs prostate glandなど取り除かれてい
る状態で、以前に、解剖される
Prostates racemose in 17.22 each ducting to penis within copulatory pouch that is confined to body
wall with accessory secretory diverticulum opening anterior to male pore and associated with a
large penial seta (as shown in Michaelsen’s fig. 1). These di
Prostates racemose in 17.22 accessory secretory diverticulum始まりがmale poreに前方であって大きい
penial setaで関連していた状態で体壁に限定されるcopulatory pouchの中の陰茎への各ダクト
(Michaelsenの図1で示されるように。) これらのdiverticulaはflattened tubular prostatesに類似して、
segments 16 or 17に達する
the current type has large flat pads around the spermathecal pores, possibly these have some
corollary function to the diverticula and penial setae of the male pores.
the current typeがthe spermathecal poresの周りに大きいflat padsを持っているということであ
る、ことによるとこれらはthe male poresのthe diverticulaとpenial setaeに何らかの
Previously thought to resemble parts of the M. hilgendorfi speciesgroup (Blakemore 2003, 2010), it is now clearly separated.
以前M. hilgendorfi species-group(Blakemore 2003、2010)の部分に類似するよ
うに思われて、それは現在、明確に切り離される。
Previously thought to resemble parts of the M. hilgendorfi speciesgroup (Blakemore 2003, 2010), it is now clearly separated.
Manus by Dr. Blakemore
A New Earthworm Genus (Oligochaeta, Megadrili, Megascolecidae) from Japan
Robert J. Blakemore1
Abstract Genus Manus is newly proposed with type and only species Pheretima koellikeri
Michaelsen, 1928. Type specimens of the type species preserved in the Zoological Museum Hamburg
are inspected and sketched. Distribution of the species is Japan (Kyushu?) with an unconfirmed Korean
record. The new genus is diagnosed by a secretory diverticulum opening adjacent to each prostate
gland and associated with a penial seta internally and, secondarily, on its manicate intestinal caeca.
Key words: Pheretima, new genus, Asiatic, prostates, conservation status, von Kölliker.
INTRODUCTION: This paper aims to establish it firmly within
bounds of ICZN (http://iczn.org/) stated aims for standard, sense
and stability for animal names in science.
manus1 -ūs, f
1 （人間の）手．
ad manum esse (Liv) 手近にある
per manūs tradere (Caes) 手渡しする
ad meam manum (Cic) 手ずから, 自ら
羅和辞典から
Appendix slide

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