65 南太平洋海域調査研究報告 No.56（2015年3月） OCCASIONAL PAPERS No.56（March 2015） ࣑ࢡࣟࢿࢩ㐃㑥࣏ࣥ࣌ᓥ࠾ࡅࡿ ₻㛫ᖏ⏕ᜥࡍࡿᕳ㈅࢟ࣂ࣐࢞ Nerita plicata ࡢ ⏕ᜥ⎔ቃẆⰍከᵝᛶ㛵ࡍࡿ◊✲ Ἑྜ Shell Color Polymorphism of Intertidal Gastropod Nerita plicata in Pohnpei Island, the Federated States of Micronesia KAWAI Kei 㮵ඣᓥᏛᅜ㝿ᓥᕋᩍ⫱◊✲ࢭࣥࢱ࣮ Research Center for the Pacific Islands, Kagoshima University せ᪨ ࢟ࣂ࣐࢞ࡣࣥࢻኴᖹὒࡢ₻㛫ᖏୖ㒊ࡢᒾ♋ᇦᗈࡃศᕸࡍࡿᕳ㈅࡛ࠊࡇࡢ௰㛫ࡣⓑⰍࡢẆࢆᣢࡘ ࡶࡢẆ⾲㯮Ⅼࢆ♧ࡍࢱࣉࡀ࠶ࡿࡇࡀሗ࿌ࡉࢀ࡚࠸ࡿࠋࡇࡢㄪᰝ࡛ࡣ࣑ࢡࣟࢿࢩ㐃㑥࣏ࣥ࣌ᕞࡢ ࣏ࣥ࣌ᮏᓥ㏆ࡃࡢᓥ࠾࠸࡚ 2014 ᖺ 8 ᭶㔝እほᐹࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊ࣏࡛ࣥ࣌ほᐹࡉࢀࡓ࢟ࣂ ࣐࢞ࡣ 3 ࡘࡢẆⰍ㸦ⓑⰍࠊ୍㒊⦤ᶍᵝࠊ㒊⦤ᶍᵝ㸧ࡢࢱࣉࡀほᐹࡉࢀࡓࠋࡇࡢከᵝᛶᙳ㡪ࡍࡿ ࡶࡢࡋ࡚㑇ఏⓗ࡞せᅉ⎔ቃⓗ࡞せᅉࡀ⪃࠼ࡽࢀࡿࠋ ἢᓊᇦࡢᒾ♋ᇦࡣ ᗘኚࡀ⃭ࡋ࠸ࡀࠊከࡃࡢ⏕≀ࡀࡑࡢ⎔ቃ㐺ᛂࡋ⏕ᜥࢆࡋ࡚࠸ࡿࠋ ࢟ࣂ࣐࢞ Nerita plicata ࡣࣥࢻኴᖹὒࡢ₻㛫ᖏୖ㒊ࡢᒾ♋ᇦᗈࡃศᕸࡍࡿᕳ㈅࡛࠶ࡿ ࠋࡇࡢ௰㛫ࡣⓑⰍࡢẆࢆᣢࡘࡶࡢẆ⾲㯮Ⅼࢆ♧ࡍࢱࣉࡀ࠶ 㸦࣎ࢵࢺ࣭ࢲࣥࢫ 1985㸧 ࡿࡇࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦࣎ࢵࢺ࣭ࢲࣥࢫ 1985, NEVILLE 2003㸧 ࠋ࢟ࣂ࣐࢞⏕ᜥࡍࡿ ᇶ㉁ࡢⰍࡢ㛵ಀࢆゎ᫂ࡍࡿࡓࡵࠊ࣑ࢡࣟࢿࢩ㐃㑥࣏ࣥ࣌ᕞࡢ࣏ࣥ࣌ᮏᓥす㒊 ⨨ࡍࡿ㏆ࡃࡢᓥ࠾ࡼࡧᓥ㛫ぢࡽࢀࡿᆺࡢᒾ࠾࠸࡚ 2014 ᖺ 8 ᭶㔝እほᐹࢆ⾜ࡗࡓࠋ ࢟ࣂ࣐࢞ࡢ㈅Ẇࡑࡢ⏕ᜥᇶ㉁ࢆ୍⥴┿ࡾࠊ◊✲ᐊᣢࡕᐤࡾࠊẆࡢከᵝᛶ ࡑࡢᇶ㉁ࡢ㛵ಀࡘ࠸࡚ㄪᰝࢆ⾜ࡗࡓࠋ 66 KAWAI Kei ᅗ 1 ほᐹࡉࢀࡓ⦤ࡀẆయぢࡽࢀࡿࢱࣉ㸦a㸧ⓑⰍࡢࢱࣉ㸦b㸧 ࣏࡛ࣥ࣌ほᐹࡉࢀࡓ࢟ࣂ࣐࢞ࡣ 3 ࡘࡢẆⰍ㸦ⓑⰍࠊ୍㒊⦤ᶍᵝࠊ㒊⦤ᶍᵝ㸧 ࡢࢱࣉࡀほᐹࡉࢀࡓ㸦ᅗ 1㸧 ࠋ୍⯡ἢᓊᇦ࡛ࡣᇶ㉁ࡢⰍࡣᵝࠎ࡛࠶ࡿࡀࠊẆࡢⰍࡣⓑⰍ ಶయࡀከࡃࢆ༨ࡵ࡚࠸ࡓࠋ⌧Ⅼ࡛ࡣࠊᇶ㉁㈅Ẇࡢከᵝᛶࡢ㛵ಀࡘ࠸࡚ࡣ᫂☜࡞ഴ ྥࡣ♧ࡉࢀ࡚࠸࡞࠸ࠋ2010 ᖺࡢ࣏ࣥ࣌ᓥ࡛ࡢㄪᰝ࠾࠸࡚ࡶ 3 ࢱࣉࡢẆከᵝᛶࡀほᐹ ࡉࢀࠊྠᵝࡢഴྥࡀほᐹࡉࢀࡓࠋࡑࡋ࡚ࠊ2010 ᖺ࡛ࡣᮏᓥࡢ㏆ࡃࡢᑠᓥ࠶ࡿሙᡤ࡛ࡣ ᇦ࡛㯮Ⰽࡢᒾࡀほᐹࡉࢀࠊࡑࡇࡢ࢟ࣂ࣐࢞ࡢẆⰍࡣ୍㒊⦤ᶍᵝࠊ㒊⦤ᶍᵝࡀࢇ ࡛ࠊⓑⰍࡢ㈅Ẇಶయࡣࢇほᐹࡉࢀ࡞ࡗࡓࠋᅇࡢㄪᰝ࡛ࡣࡇࡢࡼ࠺࡞ഴྥࡣほ ᐹࡉࢀࡎࠊఱ➼ࡢせᅉࡀẆⰍࡁࡃᙳ㡪ࢆ࠼࡚࠸ࡿࡇࡀ♧ࡉࢀࡿࠋ ࢟ࣂ࣐࢞ࡣᾋ㐟ᗂ⏕ᮇࢆᣢࡘ⪃࠼ࡽࢀࡿࡓࡵࠊ⊃࠸ᆅᇦ࡛ࡣ㑇ఏⓗྠࡌ⪃࠼ࡽ ࢀࡿࡀࠊ」㞧࡞ᆅᙧࡸ⾜ືᵝᘧ࡞ࡀ࠶ࡿẚ㍑ⓗ㏆࠸ᆅᇦ࡛ࡶ㧗࠸㑇ఏⓗከᵝᛶࡀ⥔ᣢ ࡉࢀࡿࠋᚑࡗ࡚ࠊᅇほᐹࡉࢀࡓ࣏ࣥ࣌ᮏᓥ࠾ࡅࡿẆⰍࡢ㐪࠸ࡣࠊఱࡽࡢ⎔ቃせᅉ ㉳ᅉࡋࡓ㑅ᢥᅽࡀ⛶㈅ᮇᙳ㡪ࡋࡇࡢᵝ࡞ẆⰍࡢኚ␗ࢆᘬࡁ㉳ࡇࡋࡓྍ⬟ᛶ㸦e.g., MANRÍQUEZ et al. 2009㸧 ࠊ࠶ࡿ࠸ࡣ」㞧࡞㑇ఏᏊὶືࡀࡇࡢᆅᇦࡣ࠶ࡾࠊࡇࡢせᅉࡀᘬࡁ㉳ ࡇࡋࡓྍ⬟ᛶ㸦e.g., GUERRA-VARELA et al. 2009㸧ࡢ୧᪉ࡀ⪃࠼ࡽࢀࡿࠋ ᘬ⏝ᩥ⊩ ࣎ࢵࢺ R. T.࣭ࢲࣥࢫ S. P.㸦Ἴ㒊ᛅ㔜࣭ዟ㇂ႛྖ┘ヂ㸧 1995㸬ୡ⏺ᾏ⏘㈅㢮ᅗ㚷㸪443 㡫㸪ᖹซ♫㸪ᮾி㸬 GUERRA-VARELA, J., COLSON, I., BACKELJAU, T., BREUGELMANS, K., HUGHES, R. N. and ROLAN-ALVAREZ, E. 2009. The Evolutionary Mechanism Maintaining Shell Shape and Molecular Differentiation between Two Ecotypes of the Dogwhelk Nucella lapillus. Evolutionary Ecology, 23: 261-280. MANRÍQUEZ, P. H., LAGOS, N. A., JARA, M. E. and CASTILLA, J. C. 2009. Adaptive Shell Colour Plasticity during the Early Ontogeny of an Intertidal Keystone Snail. Proceedings of the National Academy of Sciences, 106: 16298-16303. NEVILLE, C. 2003. 2002 Sea Shells: Catalogue of Indo-Pacific Mollusca, 144 pp., Neville Coleman’s Underwater Geographic Pty Ltd, Australia.
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