Ņõ'ů&k 7Ôƃ& ! mn ƈĢ½ ĀrÊą¼¿¼¿ż IJzĞÊ¿İĶĴ [email protected] Ćœ IJz'·Ī("7%õÙ;71&;Łê7#(Ɓ"($'4 %[AQIZ&4!ů!'9üŢò"(Ņő'ů& -Ôƃ& !¢6i7s(Ƃʼn&Æ!Ņ;k 7#$'4&âÝĐÂ;!79ĭæ' Ϧ&Û!ĎŽħ&Ņ'Ò;ė!728%7(7ź;¹&!ĭ æ'Ϧźwi"8)Ņ;k %źwj"7#Ņ;k 728% !'4%âÝĐÂ'ů&Ôƃ;k 7#ň 587"üİĶ"( adaptive dynamics &47¤ŊÞŘā3ůFY^cgF_e;ġ!$'4%J>V'Ņ'¡ÛŹñ 'ů;'Řā;őIJzRLOdgBćŮ'Ôƃ2š+ĿĂ(wj'ŭ6"7 ƉƌƊ ă½\Nb"(7ź;¹&!ĭæ'Ϧźwi"8)Ņ;k %źwj "7#Ņ;k 7¸¦&(ϦƇ*#ůů87#: ƉƍƊÁ Ė¦\Nb"(Ņû&ěđ&!(ĭæ'Ϧ&Û!ĎŽħ&Ņ'Ò;ė 7#ϦƇ&ů7#;ı Strict or graduated punishment? Effect of punishment strictness on the evolution of cooperation. Mayuko Nakamaru Department of Value and Decision Science, Tokyo Institute of Technology Abstract Whether or not costly punishment promotes the evolution of cooperation has been investigated not only in evolutionary biology but also in other research fields such as mathematical sciences and social sciences. The previous theoretical studies assume that each player chooses either cooperation or defection and either punishment or non-punishment; one reason is to make a simple mathematical model. In reality, different players have different cooperation level and decide how to punish others according to their cooperation level. For example, some players punish opponents strictly if their cooperation level is lower than the threshold and they do not, otherwise. Some players impose gradual punishment proportional to the opponent's cooperation level on the opponent. The former is called strict punishment and the latter is called graduate punishment. We investigate which punishment promotes the evolution of cooperation level in the complete mixing population or in the lattice structured population, by means of adaptive dynamics and computer simulations. The results are (1) strict punishment promotes the evolution of cooperation level and both coevolve in the lattice structured population, and (2) graduate punishment promotes the evolution of cooperation level and both coevolve in the complete mixing population under the condition that punishment helps cooperators invade the population occupied by defectors. ů7'9Ƒ )ãá&Įī ťŢ7[7]EHOfWRU=LO'/ IJz'·Ī("7®s'Gce ;ň 7#ō&Æ!ĩ/;s; XDgZ3ŦDgZı4& ŜŚ7'(øŸ3ŵŷ'ð#%71& %õÙ;71&;Łê7 Ŝ 7#;%ó #(Ɓ%s(;7 ;!7#ēĄţċ!. '98(ůğä¿'ŗĘ52 "Ü6'ãá&.!øŸħfŵŷħE ¼%¬Ƅ"6v'ōĚĴ¿3IJzĴ¿ HO#!2Ŝ 7'"7 &!2è:87¿žħ%İĶMgX" 'ãáĩ/;ŻÕ;(!7# 7'ů;ů71'[AQ 'Ĉ%ãá#Ņ'Ź&Įī!Ņ IZ#!ŐłĕďņĕďĬîpßÞ 'ů;ì 7#'2h ŸîpßÞRLOdgB5'ÔƃŅő ĺĭp&ĞŢfÃƆfÃŝİĶ;U=g v'ŗĘ5'İĶ27ʼnƂ PSLB%5İĶő:8!7[17, ʼn&Æ!Ņ;ő#2'"72 18] ŅÃőʼnʼn"7%5v'ʼn üŢò"(Ņő#ő'ů& #Ź#%6ŅÃő&7EHO; !èl'm;Ŕ!7#Űŋʼn& 'õĒ9 ĵ¶17#ÿ7'8&4! Æ7ŅÃő8!7ÎŇ'Ņő( #Ņů7#ň 587" à;Ĕ6[11]Ń6Ŭ®s'Gc Ĉ%ĞŢfÃƆİĶő:8![1, 23, eXDgZ&!,Ŭ;6[2] 21, 6, 3]IJzRLOdgBćŮ3åģùô ý&7#[15]©.87"( bgbŅ#'ů&Ôƃ!7 Ņ;k 5872k 72EHO #2ı8[4, 12, 13] 7ĜŇ'Ņ;ň 7'¸¦Ņ8 ĞŢħ&( Ƃʼn&Ņ;7Ƃʼn EHO;Œ71ŅōŪ(ů% ʼn*'ŅƉ¡IJzħ%ŅƊ#åģ; "($'4%[AQIZŅ'ů; ň 7#2ÿ7'i"#Ƃ ±ƌ Ņ&Ź7âÝĐÂŹñ ʼn*'Ņő'ů$'4%þy& &Ź7İĶ¤Ŋ#%7' 4!¤Ŋ$'İĶ2ů<"7[19, "üŢò"(Ĺʼn'İĶ"7Ņ' 9]ƌƐA²"'ÃƆ&4!civic âÝĐÂ&Ź7ůDgZİĶ;Ľt cooperation '|²"(Ƃʼn*'Ņ 7[14, 20] & !¡IJzħŅŗÅ87civic cooperation 'Ƈ²"(¡IJzħŅ( .6ŗÅ8%#İĶ27[8] .Ï#!Ņ;őĉćƉŤÅ%$Ɗ Nakamaru and Dieckmann (2009)'\ ;s(6i!#ŗĘ'İĶ2 Nb;Ľt7[14] Vcg]gi (Ï ů<"7[22] ¦Ɖxi; xi ! 0ƊŅ'ÒÏƉfi; fi ! 0ƊŅ' ĞŢ'őİĶ"(7Ƌ% ó'źƉui; ui ! 0Ɗ!Ņ¡ÛŹñƉai; Ņ7ƋŅ%#åģ;x ai ! 0Ɗ'Ə 8(ñ¿ħ&¢6èÄ÷%7# "(ķŸćŮ7¸¦Ɖă½\NbƊ#ķ [aLO7hõ"s(7f ŸćŮ%¸¦ƉÁĖ¦\NbƊ'ƍ %"(%Ϧ7.Ņ 'IJzRLOdgBćŮ;xÂ!7. & !2Ņ;k 7¸¦28) "(%Ņ;k 7#27! Ņ'k õ(ĭæ'Ϧ&¾! 79 )ĭæ'Ϧ&Û !ĎŽħ&Ņ'Ò;ė!72 'ůÓŨ;ê 'İĶ (ķŸćŮ'7\Nb;&!\N b;Šö7ă½\Nbi'¥ă½Ęi& Vcg]gƌs7¸¦#Vcg]g %ķă½Ę;ň 7Vcg]g(úſ îă½Ęi'Vcg]g#IJzħĭpġ 8%7(7ź;¹&! ;ő .6Vcg]gi ( xi 'Ï ĭæ'Ϧźwi"8)Ņ; ¦"Vcg]gj &7'# %źwj"7#Ņ;k EHO( C1 (xi ) = ! 1 xi" 1 #7Ċ&Vc 728% .6Ϧ;Ç g]gj (Vcg]gi 'Ϧ&Û 7#"Ņ'k õ& !Vcg]gi &Ņ;7% !'âÝĐ ±ƍ Nakamaru and Dieckmann (2009) 'ĿĂ 7¸¦($'5'Ņ;ĴâÝĐ Fj (xi ) = F "7'\Nb"( ;ő'âÝĐÂ(wj'Źñµ ĭpġ'ĭæ5'Ϧ'Í´ ( Fj (xi ) = f j exp # !1" xi u j $ ) aj % & (1) &4!ºčĝĐ.6ſî'ķă½& 'ºčĝ&Ø!½$2;Ġ0#7½$ &Ø#7Ɖ±ƌƊ'Źñµ&ůÓ 2(·üħ&(Ŗ'ÓŨ;£ŀĸĚ Ũ©.87a '&4!ŹñµĐ. »ĥ&4!Ŗ#(ĥ%7ÓŨ;£ŀ 7±ƌa ( a = 0 'ø .6Ϧ #7¥ }&EHOŅÃőEHO &45&hÖ&Ņ;k 7#âÝĐ ĭæ5'Ņ&47EHO(¥ }'Čq Â;1±ƌb (ĎŽħŅ;ıƉa = 1Ɗ ĝ&Ôƃ7#7 ±ƌƒƓ(HMLVJ>V'ŅŹñ;ı ÁĖ¦\Nb"(`eKZ&z Ɖa = 10, 1000Ɗ Fj (xi ) Vcg]gi Œ ƍs'Vcg]giř'IJzħĭpġ 7ŅEHO#%7Ņ;ÃőVcg]g ;ő½À2`eKZ&Ŕ j &7EHO( C2 ( f ) = ! 1F " 1 #7 &Ġ0#7 ķă½Ę '\Nb;ŘāÁĖ¦\Nb" Ũ#%77#ſîVcg]g'Ï ( adaptive dyamics ;ġ!ñĞ\Nb ¦ō'Ņó'ź46Ƈ#Ņ; ;ő[5]#Ņ'ůũ5% k %'"EHO25%ź #;ıķŸćŮ'7\Nb(@ 46È"2|"7#Ņ;k gG?eOWgHFY^cgF_e;ő 7#&%78&4!ů Ĉ%İĶ5ķŸćŮ&4!Ï 87#ň 587hõĎŽħ%Ņ"( ¦(ů7#:!7 .6 Ϧ;j#!2Ņ5'K[ Ņ'Ôƃ&4!ϦƇ% gG(±ƌa '4&ĸĚ¼Ƈ%7 '8#2ķŸćŮ&4!Ϧ #(%1$<$<Ϧ;j Ƈ%';7Úœ7± 7õ¨&ů0#ň 587 ƍa (Ϧ'/ůÓŨ"7# ĎŽħ%Ņ#ů7#(% 'EHO'T`[gJƉ"1Ɗ#Ï '"9Ƒ"Shimao and ¦'Ź;ı!7"1 < 1 '# Nakamaru (2013)"(\Nb'śÂ;» .6ϦƇ%!2EHO !Řā;ő[20] .65%ěđ&!ķŸćŮ'Ô ƃ&4!Ϧ(Ƈ&ů7 #:7±ƍb (EHO#ŅEH O'T`[gJƉ"1, "2Ɗ#Ə 'ůÓŨ' Shimao and Nakamaru (2013)'\Nb Ź;ı!7'ĿĂ46"1 < 1 ' ;Šö7[20]Vcg]gi (Ϧ #&2Ϧ(Ƈ&%! Ɖxi;1 ! xi ! 0ƊŅ'ÒÏƉfi; 1 ! fi ! 0ƊŅ Ņ'ÒÏƉfƊŅ'ó'źƉuƊ! 'ó'źƉui; 1 ! ui ! 0Ɗ!Ņ¡ÛŹ Ņ¡ÛŹñƉaƊ(|'.."7"1 = ñƉai; ai ! 0Ɗ'Ə 1 ×"6 "2 < 1 "(ϦhĤ 'İĶ"2ķŸćŮ7¸¦Ɖă½\NbƊ Ƈ&ů'#Ņ'ÒÏƉfƊŅ #ķŸćŮ'%¸¦ƉÁĖ¦\NbƊ' 'ó'źƉuƊ!Ņ¡ÛŹñƉaƊ( ƍ Ƈ#%!7 .6Ņ'Ôƃ&4 7.(ķŸćŮ'7\Nb;&! !ϦƇ&ů7T`[ \Nb;Šö7ă½\Nbi'¥ă½Ę gJĻ°2¾³'#Ƈ a i&Vcg]gƌs .6HMLVJ>V'Ņ¡ÛŹñƉ±ƌaƊ 'ůÓŨ;ê 'IJzRLOdgBćŮ;xÂ! ųń8!7 #7ķă½Ę(ę¥Vcg]g(ſ &ů7' î7Ǝs#ŦDgZ;őŦD Nakamaru and Dieckmann (2009)"( gZ"(¥Vcg]g(Ə ±ƌa 'HMLVJ>V'Ņ#'ů V'[eSg#%!7¥CbgV"( Ŕ58±ƌb '4%ĎŽħŅ# Əs88¥ō'Ϧ&Û! 'ů(ğ%[14]%" Ŧ*éŧ;őéŧƅ'ª&½ĝ r 9ķŸćŮ'7\Nb"(ſî ;o;´ĺų7¥Vcg]gƉj 7Vcg]g(§4%J>V'ůÓ #7Ɗ(v'[eSgƉi #7Ɗ' 'ƏsCbg ±Ǝ Shimao and Nakamaru (2013) 'ĿĂ Ï¦;2#&Ð (1) 'âÝĐÂ&Ø Sg"ŦDgZ;őČq×'ķă½ !88'Vcg]g&Ņ;7$ Ę&(ƀ¯}5`eKZ&ű)8V ;Đ17Vcg]gj ( Fj (xi ) 'ŅEH cg]g½$2;Ġ0#7 O;ç!Vcg]gi &Ņ;! ! Fj (xi ) \NbśÂ'ļŲ( Nakamaru and 'K[gG;k 7#7ŦDgZ' Dieckmann (2009)[14]#(ŏÌĥ%7 Ù5Vcg]g'ŅEHO3K[gG 'm"ú2Ŵœ%Ę(åģ'ùôbgb; ;ÑÙ#%7'Ù&Ø! » #"7"(ÙĘ¾ğ¾ ČqĝĐ.7Čq¸¦(ķă½Ę# ĝ\Nb#åģùôbgb;ġ! %7ſî7Vcg]g`eKZ&ű 7[16, 12, 13]'åģùôbgb'·"( )8!½$2;Ġ0#7½$2(·üħ vs'Ĩ;j71&ō5EHO;Œ &(Ŗ'ÓŨ;£ŀĸĚ»ĥ&4 7#HT>Oőů3# !Ŗ#(ĥ%7ÓŨ;£ŀ#7ķŸ :!7HT>Oő(Ņő#{ ćŮ긦(`eKZ&ű)8[e !6'åģùôbgb&4!Ņő ů3#:7.ÙĘ ů;ůƉƍƊÁĖ¦\Nb"( ¾ğ¾ĝ\Nb"(ķŸćŮę!2Ņ ÙĘ¾ğ¾ĝ\Nb'4%åģùôb #ő(ů3#2: gb"(ĎŽħ%Ņ¡ÛŹñ'ů !7[12, 13] ;ů7#;ı[14, 20] "(Ņ¡ÛŹñ#'ů'İĶ"( 'ƍ ÙĘ¾ğ¾ĝ\Nb#åģùôbg ůDgZİĶ"Ē3IJzŕĻ(IJ b&4!$'4%Ņ¡ÛŹñ&ů z;Ŏ71&ŅŕÂ;! 7'9Ƒ 7!Űŋʼn'ŰŋϦ&4!Ņ; ±Ǝ46ă½\Nb"( Nakamaru and k !7'4%bgbIJz}' Dieckmann (2009)[14]#§Ĉ&HMLVJ ġ'ú¼;71'þy;íİ >V'Ņ¡ÛŹñϦ'ů; Ķ27[10] ů7#:7Ɖ±Ǝ"( r = 3 # = s'âÝĐÂ'uõ&Ź7ůİĶ( 10ƊÁĖ¦\Nb'¸¦"(ĎŽħŅ şįĴ¿3ŌijľĴ¿'ĦÉ##2&ů0 Ϧ'ů;ů7#: #ň 587ůDgZĞŢ(8&Æ !~5'ëŶ3õ¨Þ;k 7¤ŊÞ "(% Shimao and Nakamaru (2013) 79 'İĶ"( sŸ'Ņ&Ź7 "(ĎŽħŅő'ůğ'9 [20]ƑÙĘ¾ğ¾ĝ\Nb"(Ņ# 'ů(Úğ7#ï"Ş; ů14'#Ë&`eKZ&Ŧ [1] Axelrod, R., 1986. An evolutionary approach DgZ;ő[eSgĐ.71Ĉ% to norms. 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