Systema Naturae∗
c Alexey B. Shipunov
v. 6.8 (November 30, 2014)
Regnum Monera [ 7Bacillus ]
Subregnum Bacteria [ 6.8Bacillus ]*
1
Superphylum Terrabacteria [ 6.2Bacillus ] s.a.
Phylum 1. Firmicutes* [ 6Bacillus ] s.a., stat.m.
Classis 1(1). Clostridia [ 5Clostridium ] s.a.2
2(2). Negativicutes [ 5Negativicoccus ]
3(3). Bacilli [ 5Bacillus ] s.a.3
4(4). Mollicutes [ 5Mycoplasma ]4
5(5). Thermi [ 5Thermus ]5
6(6). Armatimonadetes [ 5Armatimonas ] stat.m.6
7(7). Synergistia [ 5Synergistes ]
8(8). Atribacteria [ 5Caldatribacterium ] 7
9(9). Thermotogae [ 5Thermotoga ] sed.m.
10(10). Acetotermia [ 5Acetothermum ] 8
11(11). Fervidibacteria [ 5Fervidibacter ] 9
12(12). Caldiserica [ 5Caldisericum ] i.s.10
∗
Only recent taxa of indeniable living things are included. Abbreviations and signs: sed.m.
(sedis mutabilis); stat.m. (status mutabilis): s., aut i. (superior, aut interior); i.s. (incertae sedis);
sed.p. (sedis possibilis); s.s. (sensu stricto); s.a. (sensu amplo); s.aggr. (sensu aggregato); incl.
(inclusum); excl. (exclusum); p.p. (pro parte); (z) for ICZN names; † (dagger) for names introduced
here; “quotes” for environmental groups; * (asterisks) for paraphyletic taxa.
1
Incl. “Nanobacteria” i.s. et dubitativa.
2
Incl. Thermolithobacteria stat.m., Symbiobacteria, Tissierellia, Sulfobacillus i.s., Thermaerobacter i.s.
3
Incl. Erysipelotrichi, Heliobacteria.
4
Incl. Lumbricincola.
5
Incl. Thermales.
6
= “OP10”, incl. Chthonomonadetes, Fimbriimonadia.
7
=“OP10”.
8
=“OP1”.
9
=“OctSpa1-106”.
10
=“OP5”.
1
13(13). Dictyoglomia [ 5Dictyoglomus ] sed.m.
14(14). Fusobacteria [ 5Fusobacterium ] sed.m.
Phylum 2. Actinobacteres† [ 6Actinomyces ]
Classis 1(15). Actinobacteria [ 5Actinomyces ]11
Phylum 3. Cyanobacteria [ 6Nostoc ]
Classis 1(16). Gloeobacteria [ 5Gloeobacter ]12
2(17). Oscillatorineae* [ 5Oscillatoria ] s.a.13
3(18). Nostocineae [ 5Nostoc ]
Phylum 4. Chloroflexes [ 6Chloroflexus ]
Classis 1(19).
2(20).
3(21).
4(22).
5(23).
Chloroflexi [ 5Chloroflexus ]
Thermomicrobia [ 5Thermomicrobium ]14
Anaerolineae [ 5Anaerolinea ]15
Dehalococcoidetes [ 5Dehalococcoides ]16
Ktedonobacteria [ 5Ktedonobacter ]
Superphylum Gracilicutes [ 6.2Rhodospirillum ] s.a.
Phylum 5. Patescibacteria [ 6Microgenomatus ]
Classis 1(24). Gracilibacteria [ 5Altimarinus ] 18
2(25). Parcubacteria [ 5Paceibacter ] 19
3(26). Microgenomates [ 5Microgenomatus ]
17
20
Phylum 6. Spirochaetae [ 6Spirochaeta ] sed.m.
Classis 1(27). Spirochaetes [ 5Spirochaeta ] s.a.21
Phylum 7. Elusimicrobia [ 6Elusimicrobium ] sed.m.
Classis 1(28).
2(29).
3(30).
4(31).
Elusimicrobes [ 5Elusimicrobium ]22
Aerophobetes [ 5Aerophobus ] 23
Hydrogenedentes [ 5Hydrogenedens ]24
“Poribacteria” [ 5“Poribacter” ]
11
Incl. Coriobacteriia, Rubrobacteria, Thermoleophilia, Acidimicrobia.
Gloeobacter corresponds with Aphanothece.
13
Incl. Synechococcineae, ‘Prochlorophyceae’, Acaryochloris.
14
Incl. Thermobaculum, Sphaerobacter.
15
Incl. Caldilineae, Thermoflexia, Ardenticatenia.
16
Incl. Dehalobium, Dehalogenimonas.
17
Probably incl. “TM7”, “WS6”.
18
=“GN02”.
19
=“OD1”.
20
=“OP11”.
21
Incl. Leptospira.
22
= ”Endomicrobia”, “TG1”.
23
=“CD12”.
24
=“NKB19”.
12
2
5(32). Atribacteria [ 5Caldatribacterium ]25
Phylum 8. Planctobacteria [ 6Planctomyces ]26
Classis 1(33).
2(34).
3(35).
4(36).
5(37).
Planctomycetia [ 5Planctomyces ]27
Lentisphaeria [ 5Lentisphaera ]28
Verrucomicrobiae [ 5Verrucomicrobium ]29
Chlamydiae [ 5Chlamydia ]
Omnitrophica [ 5Omnitrophus ] 30
Phylum 9. Bacteroidetes [ 6Bacteroides ] s.a.31
Classis 1(38).
2(39).
3(40).
4(41).
5(42).
6(43).
7(44).
8(45).
9(46).
Cloacimonetes [ 5Cloacimonas ]32
Fibrobacteria [ 5Fibrobacter ]
Gemmatimonadetes [ 5Gemmatimonas ] sed.m.
Latescibacteria [ 5Latescibacter ]33
Chlorobia [ 5Chlorobium ]
Marinimicrobia [ 5Marinimicrobium ]34
Bacteroidia [ 5Bacteroides ]35
Ignavibacteria [ 5Ignavibacterium ]
Salinibacteria† [ 5Salinibacter ] stat.m.s.
Phylum 10. Proteobacteria [ 6Rhodospirillum ] s.a.
Classis 1(47). Epsilonproteobacteria [ 5Campylobacter ]
2(48). Calescamantes [ 5Calescibacterium ]36
3(49). Aquificae [ 5Aquifex ]
4(50). Thermodesulfobacteria [ 5Thermodesulfobacterium ] i.s.
5(51). Deferribacteres [ 5Deferribacter ]
6(52). Chrysiogenetes [ 5Chrysiogenes ]
7(53). Acidobacteria [ 5Acidobacterium ] sed.m.37
8(54). Aminicenantes [ 5Aminicenans ]38
9(55). Nitrospirae [ 5Nitrospira ]39
10(56). Desulfobacteria [ 5Desulfobacter ] s.a.40
25
=“OP9”
= ”PVC”.
27
Incl. Phycisphaerae.
28
Incl. Oligosphaera.
29
Incl. Opitutae, Spartobacteria.
30
=“OP3”.
31
=”FCB”; incl. “TG3”.
32
=“WWE1”.
33
=“WS3”.
34
=“SAR406”; incl. Caldithrix sed.m.
35
Incl. Flavobacteria, Sphingobacteria, Cytophagia.
36
=“EM19”.
37
Incl. “TM6” stat.m., Halophagae, Solibacteres.
38
=“OP8”.
39
Incl. Nitrospinae.
40
Incl. Deltaproteobacteria.
26
3
11(57). Rhodobacteria [ 5Rhodobacter ] s.a.41
Subregnum Archaebacteria [ 6.8Methanobacterium ]*
Phylum 11. Euryarchaeota [ 6Methanobacterium ]42
Classis 1(58).
2(59).
3(60).
4(61).
5(62).
6(63).
7(64).
8(65).
9(66).
Parvarcheota [ 5Parvarchaeum ] 43
Diapherotrites [ 5Iainarchaeum ] 44
Nanoarchea [ 5Nanoarchaeum ]
Aenigmarcheota [ 5Aenigmarchaeum ] 45
Thermococci [ 5Thermococcus ]
Methanobacteria [ 5Methanobacterium ] s.a.46
Thermoplasmata [ 5Thermoplasma ]
Archaeoglobi [ 5Archaeoglobus ]
Halomebacteria [ 5Halobacterium ] 47
Phylum 12. Filarchaeota* [ 6Sulfolobus ]48
Classis 1(67).
2(68).
3(69).
4(70).
Sulfolobea [ 5Sulfolobus ]49
Thermoprotei [ 5Thermoproteus ]50
Cenarchaea [ 5Cenarchaeum ]51
Korarchaea [ 5Korarchaeum ] stat.m.
Regnum Protista [ 7Euglena ]*52
Subregnum Excavata [ 6.8Euglena ]
Phylum 13. Metamonada [ 6Trichomonas ] s.a.
Classis 1(71). Anaeromonadea [ 5Oxymonas ]53
2(72). Parabasalea [ 5Trichomonas ]54
3(73). Eopharyngea [ 5Hexamita ]55
41
Incl.
Alphaproteobacteria, Acidithiobacillia, Zetaproteobacteria, Betaproteobacteria,
Gammaproteobacteria.
42
=”DPANN”.
43
=“ARMAN” group.
44
=“pMC2A384”.
45
=“DSEG”.
46
Incl. Methanobacteriales, Methanococcales stat.m., Methanopyrales i.s.
47
Incl. Nanohaloarchaea, Methanomicrobiales, Methanosrcinales, Halobacteriales.
48
=”TACK”.
49
Incl. Desulfurococcales, Caldisphaerales.
50
Incl. ’Geoarchaeote NAG1’.
51
Incl. Thaumarcheota, Caldiarchaeum (“Aigarcheota”).
52
Incl. numerous unplaced genera from (see Patterson, 2000), i.e., Magosphaera.
53
Incl. Oxymonadida, Trimastix.
54
Incl. Trichomonadea
55
Incl. Carpediemonadida, Retortamonadida stat. m., Diplomonadida.
4
Phylum 14. Discoba [ 6Euglena ] s.a.
Subphylum Loukozoa [ 5.8Jacoba ]
Classis 1(74). Malawimonadea [ 5Malawimonas ] sed.m.
2(75). Tsukubea [ 5Tsukubamonas ]
3(76). Jakobea [ 5Jakoba ]56
Subphylum Percolozoa [ 5.8Naegleria ]
Classis 4(77). Heterolobosea [ 5Naegleria ]57
Subphylum Euglenozoa [ 5.8Euglena ]
Classis 5(78). Euglenophyceae [ 5Euglena ]* s.a.58
6(79). Kinetoplastea [ 5Bodo ]
Subregnum Panmycota† [ 6.8Agaricus ]*
Phylum 15. Sulcozoa [ 6Apusomonas ]
Classis 1(80).
2(81).
3(82).
4(83).
5(84).
Anisomonadea [ 5Collodictyon ] sed.m.59
Apusomonadea [ 5Apusomonas ]
Breviatea [ 5Breviata ]60
Ancyromonadea [ 5Ancyromonas ] sed.m.61
Hilomonadea sed.m.[ 5Micronuclearia ]62
Phylum 16. Amoebozoa [ 6Amoeba ]
Subphylum Lobosea [ 5.8Amoeba ]63
Classis 1(85). Tubulinea [ 5Amoeba ]64
2(86). Discosea [ 5Acanthamoeba ]
Subphylum Conosea [ 5.8Physarum ]
Classis 3(87). Archamoebae [ 5Mastigamoeba ]65
4(88). Variosea [ 5Filamoeba ]* sed.m.66
5(89). Macromycetozoa [ 5Physarum ]67
Phylum 17. Ophistosporidia [ 6Rozella ]*
Classis 1(90). Cristidiscoidea [ 5Nuclearia ]
56
68
Incl. Andalucia
Incl. Acrasis, Percolomonas, Pharyngomonas, Stephanopogon, etc.
58
Incl. Diplonemea (= Diplosonematea) stat.m., Symbiontida, Bihospites, Postgaardi, Notosolenus, Petalomonas.
59
Incl. Diphylleia (z), Collodictyon, Sulcomonas.
60
Incl. Subulatomonas, Pygsuia.
61
Discocelia (Discocelis) sed.m.
62
Incl. Micronuclearia, Rigifila.
63
Incl. Corallomyxa i.s, Stereomyxa i.s.
64
Incl. Copromyxa.
65
Incl. Rhizomastix
66
Incl. Protostelia p.p., Multicilia, Phalansterium, Ceratiomyxella, Acramoeba etc.
67
Ceratiomyxa, Dictyostelia, Myxogastria, Protostelia p.p., Semimorula.
68
Incl. Fonticula.
57
5
2(91). Aphelidea [ 5Aphelidium ]
3(92). Cryptomycetes [ 5Rozella ]69
4(93). Microsporea [ 5Microsporidium ]70
Phylum 18. Eomycota† [ 6Mucor ]* s.a.
Classis 1(94).
2(95).
3(96).
4(97).
5(98).
6(99).
Chytridiomycetes [ 5Chytridiopsis ]71
Blastocladiomycetes [ 5Blastocladium ]
Olpidiomycetes [ 5Olpidium ]
Kickxellomycetes [ 5Kickxella ]72
Mucoromycetes [ 5Mucor ]*73
Glomeromycetes [ 5Glomus ]74
Phylum 19. Basidiomycota [ 6Agaricus ]
Subphylum Ustilagomycotina [ 5.8Ustilago ]
Classis 1(100). Exobasidiomycetes [ 5Exobasidium ]
2(101). Ustilaginomycetes [ 5Ustilago ]
Subphylum Pucciniomycotina [ 5.8Uredo ]
Classis 3(102). Pucciniomycetes [ 5Uredo ]
4(103). Atractiellomycetes [ 5Atractiella ]
5(104). Cystobasidiomycetes [ 5Cystobasidium ]
6(105). Agaricostilbomycetes [ 5Agaricostilbum ]
7(106). Microbotryomycetes [ 5Microbotryum ]75
8(107). Mixiomycetes [ 5Mixia ]
Subphylum Agaricomycotina [ 5.8Agaricus ]
Classis 9(108). Wallemiomycetes [ 5Wallemia ] 76
10(109). Bartheletiomycetes† [ 5Bartheletia ]
11(110). Tremellomycetes [ 5Tremella ]
12(111). Dacrymycetes [ 5Dacrymyces ]
13(112). Agaricomycetes [ 5Agaricus ]
Phylum 20. Ascomycota [ 6Ascomyces ]77
Subphylum Taphrinomycotina [ 5.8Ascomyces ]*78
Classis 1(113). Neolectomycetes [ 5Neolecta ]
2(114). Taphrinomycetes [ 5Ascomyces ]79
69
Incl.
Incl.
71
Incl.
72
Incl.
73
Incl.
74
Incl.
75
Incl.
76
Incl.
77
Incl.
78
Incl.
79
Incl.
70
Caulochytrium
Mitosporidium, Paramicrosporidium, Metchnikovellidea sed.m., Nucleophaga
Monoblepharidales incl. Helicosporidium, Neocallimastigales, Thalassochytrium sed.m.
Harpellales, Asellariales, Zoopagales, Entomophthorales, Nephridiophaga.
Endogonales, Mortierellales stat.m.
Glomeromycetes, Archaeosporomycetes, Paraglomeromycetes.
Cryptomycocolacales, Classiculales.
Basidioascus, Geminibasidium, Entorrhiza stat.m.i.
Spirogyromyces i.s.
“SCGI” group.
Saitoella, Schizosaccharomycetes, Pneumocystis.
6
3(115). Saccharomycetes [ 5Saccaromyces ]
4(116). Archaeorhizomycetes [ 5Archaeorhizomyces ]
Subphylum Pezizomycotina [ 5.8Tuber ]
Classis 5(117). Pezizomycetes [ 5Tuber ]
6(118). Orbiliomycetes [ 5Orbilia ]
7(119). Laboulbeniomycetes [ 5Laboulbenia ]
8(120). Xylonomycetes [ 5Xylona ]
9(121). Dothideomycetes [ 5Dothidea ]80
10(122). Leotiomycetes [ 5Leotia ]81
11(123). Sordariomycetes [ 5Sordaria ]
12(124). Geoglossomycetes [ 5Geoglossum ]
13(125). Lichinomycetes [ 5Lichina ]82
14(126). Eurotiomycetes [ 5Penicillium ]
15(127). Lecanoromycetes [ 5Lecanora ]83
Phylum 21. Choanozoa [ 6Codosiga ] s.a.
Classis 1(128). Filasterea [ 5Capsaspora ]84
2(129). Choanomonadea [ 5Codosiga ]85
3(130). Ichthyosporea [ 5Ichthyophonus ]86
Subregnum Panalgae† [ 6.8Fucus ]
Infraregnum Chromobionta [ 6.5Fucus ] s.a.87
Superphylum Rhizaria [ 6.2Cercomonas ]
Phylum 22. Cercozoa [ 6Cercomonas ] s.a.88
Subphylum Filosea [ 5.8Cercomonas ]
Classis 1(131). Hemimastigea [ 5Spironematella ] i.s.89
2(132). Ventrifilosea [ 5Coelodendrum ]* s.a.90
3(133). Sarcomonadea [ 5Cercomonas ] s.a91
4(134). Metromonadea [ 5Metromonas ] s.a.92
80
Incl. Arthoniales, Symbiotaphrinales sed.m.
Incl. Trizobia.
82
Incl. Coniocybomycetes.
83
Incl. Candelariales sed.m.
84
Incl. Ministeria.
85
Incl. Corallochytrium.
86
Incl. Amoebidium, Eccrinales.
87
= Sar.
88
Incl. Meteora i.s.
89
Incl. Hemimastix, Stereonema, Paramastix sed.m.
90
Incl. Phaeodarea, Ebriida, Cryomonadida, Spongomonadidae, Thaumatomonadidae, Euglyphida, Luffisphaera Paraluffisphaera, Rotosphaerida, Pompholyxophrys, Mataza, Chlamydophryidae.
91
Cercomonadida, Pansomonadida, Glissomonadida; incl. Guttulinopsis.
92
Incl. Metopion etc.
81
7
5(135). Granofilosea [ 5Heliomorpha ]93
6(136). Skiomonadea [ 5Tremula ]
7(137). Chlorarachniophyceae [ 5Chlorarachnion ]
Subphylum Endomyxa [ 5.8Gromia ]
Classis 8(138). Gromiidea [ 5Gromia ]
9(139). Ascetosporea [ 5Haplosporidium ]94
10(140). Proteomyxidea [ 5Filoreta ]*95
11(141). Plasmodiophorea [ 5Plasmodiophora ]96
Phylum 23. Retaria [ 6Acanthometra ]
Classis 1(142).
2(143).
3(144).
4(145).
5(146).
6(147).
Polycystinea [ 5Collosphaera ]*
Taxopodida [ 5Sticholonche ] stat.m.
Acantharia [ 5Acanthometra ]
Monothalamea [ 5Allogromia ]98
Tubothalamea [ 5Miliolina ]
Globothalamea [ 5Rotalia ]
97
Superphylum Heterokonta [ 6.2Fucus ]
Phylum 24. Labyrinthomorpha [ 6Labyrinthula ]
Classis 1(148). Labyrinthulea [ 5Labyrinthula ]99
Phylum 25. Opalozoa† [ 6Opalina ]
Classis 1(149).
2(150).
3(151).
4(152).
5(153).
Bicoecea [ 5Bicosoeca ]
Blastocystea [ 5Blastocystis ]
Placididea [ 5Placidia ]100
Nanomonadea [ 5Solenicola ]101
Opalinea [ 5Opalina ]102
Phylum 26. Oomycota [ 6Saprolegnia ]
Classis 1(154). Oomycetes [ 5Saprolegnia ]103
93
Incl. Massisteria, Desmothoracida, Gymnosphaeridae sed.m., Heliomonadida, Leucodictyon,
Reticuloamoeba, etc.
94
Incl. Paramyxidia, Bonamia, Claustrosporidium, Paradinium, Mikrocytos mackini i.s.
95
Incl. Vampyrellidae stat.m., Biomyxidae, Pseudospora sed.m., Rhizoplasmidae s.aggr., Reticuloamoeba sed.m.
96
Incl. Phagomyxa.
97
Incl. Larcopyle.
98
Incl. Xenophyophorea, Reticulomyxidae, Komokiacea, Schizocladus.
99
Excl. Labyrinthomyxa.
100
Incl. Wobbia, Rictus sed.m.
101
Incisomonas, “MAST-3”.
102
Incl. Proteromonadida.
103
Incl. Pirsonia, Hyphochitriomycetales, Developayella.
8
Phylum 27. Chromophyta [ 6Fucus ]
Subphylum Pelagophytina [ 5.8Pelagomonas ]
Classis 1(155). Bacillariophyceae [ 5Diatoma ] s.a.
2(156). Bolidophyceae [ 5Bolidomonas ] 104
3(157). Dictyochophyceae [ 5Dictyocha ] 105
4(158). Pelagophyceae [ 5Pelagomonas ] 106
5(159). Pinguiophyceae [ 5Pinguiochrysis ]
Subphylum Chrysophytina [ 5.8Chrysococcus ]
Classis 6(160). Eustigmatophyceae [ 5Eustigmatos ]
7(161). Chrysophyceae [ 5Chrysococcus ] s.a.107
Subphylum Xanthophytina [ 5.8Tribonema ]
Classis 8(162). Xanthophyceae [ 5Tribonema ]108
9(163). Phaeophyceae [ 5Fucus ] s.a.109
10(164). Raphidophyceae [ 5Rhaphidomonas ]110
11(165). Actinophryida [ 5Actinophrys ]
Superphylum Alveolata [ 6.2Paramecium ]
Phylum 28. Myzozoa [ 6Peridinium ]
Classis 1(166). Protalveolata [ 5Colponema ]
2(167). Colponemida [ 5Colponema ]111
Subphylum Dinozoa [ 5.8Peridinium ]
Classis 3(168). Perkinsida [ 5Perkinsus ]112
4(169). Squirmidea [ 5Filipodium ]113
5(170). Psammosea [ 5Psammosa ]
6(171). Ellobiopsea [ 5Ellobiopsis ]
7(172). Syndinea [ 5Syndinium ]114
8(173). Oxyrridea [ 5Oxyrris ]
9(174). Dinoflagellata [ 5Peridinium ]115
Subphylum Apicomplexa [ 5.8Plasmodium ]116
Classis 10(175). Chromerida [ 5Chromera ]117
104
Incl.
Incl.
106
Incl.
107
Incl.
108
Incl.
109
Incl.
110
Incl.
111
Incl.
112
Incl.
113
Incl.
114
Incl.
115
Incl.
116
Incl.
117
Incl.
105
Parmales.
Pedinellales, Rhizochromulinales.
Sarcinochrysidales, Olisthodiscus sed.m.
Picophagus, Synchroma, Leukarachnion, Paraphysomonas.
Chrysomeridophyceae stat.m.
Schizocladiophyceae, Phaeothamniales, Aurearenophyceae.
Olisthodiscus sed.m.
Acavomonas stat.m.
Perkinsus, Parvilucifera, Rastrimonas, Phagodinium sed.m.
Platyproteum.
“Marine Alveolate Groups I and II”.
Noctiluca stat.m.
‘X-cells’ i.s.
Vitrella
9
11(176).
12(177).
13(178).
14(179).
Apicomonadea [ 5Colpodella ]
Paragregarea [ 5Selenidium ]
Gregarinomorphea [ 5Gregarina ]118
Coccidiomorphea [ 5Plasmodium ] 119
Phylum 29. Ciliophora [ 6Paramecium ]
Subphylum Postciliodesmatophora [ 5.8Spirostomum ]
Classis 1(180). Karyorelictea [ 5Trachelocera ]
2(181). Heterotrichea [ 5Spirostomum ]120
Subphylum Ventrata [ 5.8Paramecium ]121
Classis 3(182). Phyllopharyngea [ 5Podophrya ]122
4(183). Nassophorea [ 5Nassula ]123
5(184). Colpodea [ 5Colpoda ]
6(185). Prostomatea [ 5Prorodon ]
7(186). Plagiopylea [ 5Plagiopyla ]
8(187). Oligohymenophorea [ 5Paramecium ]
Subphylum Spirotrichia [ 5.8Oxytricha ]
Classis 9(188). Litostomatea [ 5Didinium ]
10(189). Armophorea [ 5Metopus ]124
11(190). Spirotrichea [ 5Oxytricha ]
12(191). Cariacotrichea [ 5Cariacotrix ]
Subphylum Lamellicorticata [ 5.8Clevelandella ]
Classis 13(192). Armophorea [ 5Clevelandella ]
14(193). Litostomatea [ 5Spathidium ]
Infraregnum Hacrobia [ 6.5Cryptomonas ] s.m.
Phylum 30. Cryptista [ 6Cryptomonas ] s.a.
Classis 1(194).
2(195).
3(196).
4(197).
Picozoa [ 5Picomonas ]
Cryptophyceae [ 5Cryptomonas ]
Leucocrypta [ 5Katablepharis ] sed.m.125
Palpitea [ 5Palpitomonas ]
Phylum 31. Heliozoa [ 6Pterocystis ] sed.m.
Classis 1(198). Microhelea [ 5Microheliella ]
2(199). Centrohelea [ 5Pterocystis ]
118
Incl. Cryptosporidium.
Incl. Nephromyces, Piroplasmida.
120
Incl. Protocruzia sed.m.
121
= ”Conthreep”; incl. Discotrichida i.s.
122
Incl. Suctoria, Synchimeniida.
123
Incl. Nassulina, Microthoracina.
124
Incl. Licnophorida sed.m.
125
Incl. Leucocryptos, Platychilomonas, Hatena, Roombia.
119
10
Phylum 32. Haptophyta [ 6Prymnesium ]
Classis 1(200). Telonemea [ 5Telonema ] stat.m.
2(201). Prymnesiophyceae [ 5Prymnesium ]126
3(202). “Rappenomonadae” [ 5“Rappenomonas” ]127
Infraregnum Chlorobionta [ 6.5Volvox ]
Phylum 33. Glaucophyta [ 6Glaucocystis ]
Classis 1(203). Glaucophyceae [ 5Glaucocystis ]
Phylum 34. Rhodophyta [ 6Bangia ]
Classis 1(204).
2(205).
3(206).
4(207).
5(208).
Cyanidiophyceae [ 5Cyanidium ]
Rhodellophyceae s.a.[ 5Rhodella ]128
Compsogonophyceae [ 5Compsopogon ]
Bangiophyceae [ 5Bangia ]
Florideophyceae [ 5Palmaria ]
Phylum 35. Chlorophyta [ 6Volvox ]* s.a.
Subphylum Chlorophytina [ 5.8Volvox ]
Classis 1(209). Prasinococcophyceae [ 5Prasinococcus ]129
2(210). Pyraminomonadophyceae [ 5Pyraminomonas ]
3(211). Mamiellophyceae [ 5Mamiella ]
4(212). Nephroselmidophyceae [ 5Nephroselmis ]* s.a.130
5(213). Pedinophyceae [ 5Pedinomonas ] sed.m.131
6(214). Chlorodendrophyceae [ 5Tetraselmis ]132
7(215). Chlorellophyceae [ 5Chlorella ]
8(216). Trebouxiophyceae [ 5Trebouxia ]133
9(217). Chlorophyceae [ 5Volvox ]
10(218). Ulvophyceae [ 5Ulva ]
Subphylum Charophytina [ 5.8Chara ]*
Classis 11(219). Mesostigmatophyceae [ 5Mesostigma ]134
12(220). Charophyceae [ 5Chara ]*135
126
Incl.
Incl.
128
Incl.
129
Incl.
130
Incl.
131
Incl.
132
Incl.
133
Incl.
134
Incl.
135
Incl.
127
Pavlovophyceae.
“OM270”.
Stylonematophyceae, Porphyridiophyceae.
Prasinoderma, Palmophyllales (Palmophyllum, Verdigellas).
“clade VIII” Pycnococcaceae, “clade IX” Picocystis, “CCMP-1205”.
Chlorochytridion, Resultomonas, Marsupiomonas
Scherffelia, Oltmannsielliopsis sed.m.
Geminella, Helicosporidium, Prasiola.
Chlorokybus sed.m.
Klebsormidiophyceae stat.m., Conjugatophyceae.
11
Regnum Vegetabilia [ 7Magnolia ]
Phylum 36. Bryophyta [ 6Bryum ]*
Subphylum Hepaticae [ 5.8Marchantia ]
Classis 1(221). Haplomitriopsida [ 5Haplomitrium ]136
2(222). Marchantiopsida [ 5Marchantia ]137
3(223). Jungermanniopsida [ 5Jungermannia ]138
Subphylum Bryophytina [ 5.8Bryum ]
Classis 4(224). Takakiopsida [ 5Takakia ]
5(225). Sphagnopsida [ 5Sphagnum ]
6(226). Andreaeopsida [ 5Andreaea ]139
7(227). Polytrichopsida [ 5Polytrichum ]*140
8(228). Bryopsida [ 5Bryum ]141
Subphylum Anthocerotophytina [ 5.8Anthoceros ]
Classis 9(229). Anthocerotopsida [ 5Anthoceros ]142
Phylum 37. Pteridophyta [ 6Pteris ]*
Subphylum Lycopodiophytina [ 5.8Lycopodium ]
Classis 1(230). Lycopodiopsida [ 5Lycopodium ]143
Subphylum Pteridophytina [ 5.8Pteris ]
Classis 2(231). Equisetopsida [ 5Equisetum ]
3(232). Psilotopsida [ 5Psilotum ]
4(233). Ophioglossopsida [ 5Ophioglossum ]
5(234). Marattiopsida [ 5Marattia ]
6(235). Pteridopsida [ 5Pteris ]
Phylum 38. Spermatophyta [ 6Magnolia ] s.a.
Classis 1(236).
2(237).
3(238).
4(239).
5(240).
136
Incl.
Incl.
138
Incl.
139
Incl.
140
Incl.
141
Incl.
142
Incl.
143
Incl.
144
Incl.
137
Ginkgoopsida [ 5Ginkgo ]
Cycadopsida [ 5Cycas ]
Angiospermae [ 5Magnolia ]
Gnetopsida [ 5Gnetum ] stat.m.
Pinopsida [ 5Pinus ]144
Treubiales.
Blasiales, Sphaerocarpales, Monocleales.
Pelliales, Fossombroniales, Pallaviciniales.
Andreaeobryum stat.m.i.
Tetraphidales i.s, Oedipodiales stat.m.
Buxbaumiales sed.m., Diphysciales sed.m., Timmiales.
Leiosporocerotales.
Iso¨etopsida stat.m.
Cupressopsida.
12
Regnum Animalia [ 7Araneus ]145
Subregnum Ctenophora [ 6.8Ctenoplana ]
Phylum 39. Ctenophora [ 6Ctenoplana ]
Classis 1(241). Ctenophoroidea [ 5Ctenoplana ]146
Subregnum Spongia [ 6.8Spongia ]
Phylum 40. Porifera [ 6Spongia ]
Subphylum Silicea [ 5.8Spongia ] stat.m.
Classis 1(242). Hexactinellea [ 5Euplectella ]
2(243). Demospongia [ 5Spongia ] s.aggr.
Subphylum Calcispongia [ 5.8Sycetta ] stat.m.
Classis 3(244). Calcarea [ 5Sycetta ]
4(245). Homoscleromorpha [ 5Oscarella ]
Subregnum Phagocytellozoa [ 6.8Trichoplax ]
Phylum 41. Placozoa [ 6Trichoplax ]
Classis 1(246). Trichoplacoidea [ 5Trichoplax ]
Subregnum Cnidaria [ 6.8Hydra ]147
Phylum 42. Cnidaria [ 6Hydra ]
Classis 1(247).
2(248).
3(249).
4(250).
5(251).
6(252).
7(253).
8(254).
Hexacorallia [ 5Actinia ]
Octocorallia [ 5Alcyonium ]
Coronatae [ 5Linuche ]
Staurozoa [ 5Lucernaria ]
Cubozoa [ 5Carybdea ]
Scyphozoa [ 5Aurelia ]
Hydrozoa [ 5Hydra ]
Polypodiozoa [ 5Polypodium ] sed.m.
Phylum 43. Myxozoa [ 6Myxidium ]
Classis 1(255). Malacosporea [ 5Tetracapsula ]148
2(256). Myxosporea [ 5Myxidium ]149
145
Incl. Salinella i.s.
Incl. Tentaculifera, Nuda.
147
Incl. Dendrogramma i.s.
148
= Tetracapsula, Buddenbrockia.
149
= Actinomyxidia.
146
13
Subregnum Bilateria [ 6.8Araneus ]
Infraregnum Acoelomorpha [ 6.5Convoluta ]
Phylum 44. Acoelomorpha [ 6Convoluta ]
Classis 1(257). Acoela [ 5Convoluta ]
2(258). Nemertodermatida [ 5Nemertoderma ]
3(259). Xenoturbelloidea [ 5Xenoturbella ]
Infraregnum Deuterostomia [ 6.5Felis ]
Phylum 45. Echinodermata [ 6Echinus ]
Classis 1(260).
2(261).
3(262).
4(263).
5(264).
Crinoidea [ 5Metacrinus ]
Asteroidea [ 5Asterias ]150
Echinoidea [ 5Echinus ]
Ophiuroidea [ 5Ophiura ]
Holothurioidea [ 5Holothuria ]
Phylum 46. Hemichordata [ 6Balanoglossus ]
Classis 1(265). Enteropneusta [ 5Balanoglossus ]*
2(266). Pterobranchia [ 5Rhabdopleura ]
Phylum 47. Chordata [ 6Felis ] s.a.
Subphylum Cephalochordata [ 5.8Branchiostoma ]
Classis 1(267). Leptocardii [ 5Branchiostoma ]
Subphylum Vertebrata [ 5.8Felis ]
Classis 2(268). Cyclostomata [ 5Myxine ]*
3(269). Chondrichtyes [ 5Squalus ]
4(270). Actinopterygii [ 5Perca ]
5(271). Dipnoi [ 5Protopterus ]152
6(272). Amphibia [ 5Rana ]
7(273). Reptilia [ 5Gecko ]*
8(274). Aves [ 5Gallus ] stat.m.s.
9(275). Mammalia [ 5Felis ]
Subphylum Tunicata [ 5.8Ascidia ] stat.m.i.
Classis 10(276). Ascidiacea [ 5Ascidia ]153
150
Incl.
Incl.
152
Incl.
153
Incl.
151
Xyloplax.
Planctosphaera.
Latimeria.
Thaliacea, Soberacea, Larvacea stat.m.
14
151
Infraregnum Protostomia [ 6.5Araneus ]
Superphylum Spiralia [ 6.2Nereis ]
Phylum 48. Platyzoa [ 6Fasciola ] s.a.
Subphylum Gnathifera [ 5.8Rotifer ] s.a.
Classis 1(277). Gnathostomulida [ 5Gnathostomula ]154
2(278). Micrognathozoa [ 5Limnognathia ] sed.m.
3(279). Syndermata [ 5Rotifer ]155
Subphylum Gastrotricha [ 5.8Macrodasys ] stat.m.i.
Classis 4(280). Gastrotrichoidea [ 5Macrodasys ]
Subphylum Platyhelminthes [ 5.8Fasciola ]
Classis 5(281). Catenulida [ 5Catenula ]
6(282). Rhabditophora [ 5Planaria ]*156
7(283). Neodermata [ 5Fasciola ]157
Phylum 49. Mollusca [ 6Limax ]
Classis 1(284).
2(285).
3(286).
4(287).
5(288).
6(289).
7(290).
8(291).
Solenogastres [ 5Neomenia ]
Caudofoveata [ 5Chaetoderma ]
Polyplacophora [ 5Chiton ]
Monoplacophora [ 5Monoplacophorus ]
Cephalopoda [ 5Octopus ]
Scaphopoda [ 5Dentalium ]
Gastropoda [ 5Limax ]
Bivalvia [ 5Mytilus ]
Phylum 50. Lophophorata [ 6Lingula ]
Subphylum Brachiozoa [ 5.8Lingula ]
Classis 1(292). Linguliformea [ 5Lingula ]
2(293). Craniiformea [ 5Craniscus ]
3(294). Rhynchonelliformea [ 5Rhynchonella ]
4(295). Phoronida [ 5Phoronis ]
Subphylum Ectoprocta [ 5.8Flustra ]
Classis 5(296). Gymnolaemata [ 5Flustra ] 158
6(297). Phylactolaemata [ 5Plumatella ]
Subphylum Kamptozoa [ 5.8Pedicellina ] stat.m.
Classis 7(298). Cycliophora [ 5Symbion ]
8(299). Entoprocta [ 5Pedicellina ]
154
Incl.
Incl.
156
Incl.
157
Incl.
158
Incl.
155
Filospermoidea, Bursovaginoidea.
Seison, Bdelloidea, Acanthocephala s.l., Monogononta.
Macrostomida, Polycladida, Neoophora s.s.
Monogenea, Trematoda, Cestoda.
Stenolaemata.
15
Phylum 51. Nemertea [ 6Nemertes ]
Classis 1(300). Archynchocoela [ 5Archynchonemertes ]
2(301). Nemertini [ 5Nemertes ]159
Phylum 52. Annelida [ 6Nereis ]
Classis 1(302).
2(303).
3(304).
4(305).
Sipunculida [ 5Sipunculus ] s.a.160
Myzostomida [ 5Myzostoma ] sed.m.
Pleistoannelida* [ 5Nereis ]161
Clitellata [ 5Limbricus ] 162
Phylum 53. Mesozoa [ 6Dicyema ] sed.m.
Classis 1(306). Dicyemida [ 5Dicyema ]
2(307). Orthonectida [ 5Rhopalura ]
Superphylum Chaetozoa† [ 6.2Sagitta ]
Phylum 54. Chaetognatha [ 6Sagitta ]
Classis 1(308). Sagittoidea [ 5Sagitta ]
Superphylum Ecdysozoa [ 6.2Araneus ]
Phylum 55. Nemathelminthes [ 6Ascaris ] s.a.
Classis 1(309).
2(310).
3(311).
4(312).
5(313).
Priapulida [ 5Priapulus ]
Kinorhyncha [ 5Kinorhynchus ]
Loricifera [ 5Nanaloricus ]
Nematomorpha [ 5Gordius ]
Nematoda [ 5Ascaris ]163
Phylum 56. Tardigrada [ 6Macrobiotus ] stat.m.
Classis 1(314). Tardigradoidea [ 5Macrobiotus ]164
Phylum 57. Arthropoda [ 6Araneus ] s.a.
Subphylum Lobopoda [ 5.8Peripatus ] stat.m.
Classis 1(315). Onychophora [ 5Peripatus ]
Subphylum Cheliceromorpha [ 5.8Araneus ]
Classis 2(316). Pantopoda [ 5Pycnogonum ]
3(317). Chelicerata [ 5Araneus ]165
159
Incl. Anopla, Enopla.
Incl. Oweniidae stat.m., Magelonidae stat.m., Chaetopteridae stat.m., Amphinomidae stat.m.
161
Incl. Echiura, Sibolginida (Pogonophora et Vestimentifera), Lobatocerebrum sed.m., Jennaria
sed.m., Diurodrilus.
162
Incl. Aelosomata.
163
Incl. Adenophorea, Secernentea.
164
Incl. Heterotardigrada, Mesotardigrada i.s., Eutardigrada.
165
Incl. Xiphosura.
160
16
Subphylum Myriapoda [ 5.8Scolopendra ]
Classis 4(318). Chilopoda [ 5Scolopendra ]
5(319). Symphyla [ 5Symphylella ]
6(320). Pauropoda [ 5Pauropus ]
7(321). Diplopoda [ 5Julus ]
Subphylum Pancrustacea [ 5.8Scarabaeus ]
Classis 8(322). Ostracoda [ 5Cypris ]
9(323). Oligostraca [ 5Argulus ] s.a., stat.m.166
10(324). Copepodoidea [ 5Cyclops ]
11(325). Thecostraca [ 5Lepax ] 167
12(326). Malacostraca [ 5Cancer ]168
13(327). Branchiopoda [ 5Daphnia ]
14(328). Hexapoda [ 5Scarabaeus ] s.a.169
15(329). Remipedia [ 5Speleonectes ]
16(330). Cephalocarida [ 5Hutchinsoniella ]
166
Incl.
Incl.
168
Incl.
169
Incl.
167
Branchiura, Pentastomida sed.m., Mystacocarida sed.m.
Tantulocarida, Facetotecta, Ascothoracida.
Leptostraca.
Entognatha
17

航空 - JTB

IV-2. 分子進化分野 - 筑波大学計算科学研究センター