Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
[Afromorgus Scholtz, 1986]
Coleotteri Scarabeoidei d’Italia.
[Afromorgus melancholicus (Fåhraeus, 1857)]
Elenco Sistematico (luglio 2014)
LUCANIDAE
TroxFabricius, 1775
[Trox cadaverinus
Latreille, 1804
*Trox clathratusReiche, 1861
AESALINAEMacLeay, 1819
Trox cribrum clathratusReiche, 1861
AESALINI MacLeay, 1819
AesalusFabricius, 1801
Aesalus scarabaeoides
*Trox cribrum
Gené, 1836
*Trox fabricii
Reiche, 1853
[Trox granulipennisFairmaire, 1852]
(Panzer, 1795)
*Trox litoralis
*Aesalus scarabaeoides scarabaeoides(Panzer, 1795)
Pittino, 1991
*Trox nigerRossi, 1792
*Aesalus scarabaeoides meridionalisBartolozzi, 1989
Trox hipidus niger Rossi, 1792
*Aesalus scarabaeoides siculus Baviera, 200
*Trox nodulosusHarold, 1872
Trox hispidus nodulosus Harold, 1872
SYNDESINIMacLeay, 1819
*Trox perlatus
CeruchusMacLeay, 1819
*Trox perrisi
*Ceruchus chrysomelinus
Illiger, 1802]
(Goeze, 1777)
Fairmaire, 1868
*Trox sabulosus
(Hochenwart, 1785)
(Linnaeus, 1758)
*Trox scaber (Linnaeus, 1767)
SinodendronHellwig, 1792
*Sinodendron cylindricum(Linnaeus, 1758)
BOLBOCERATIDAE Mulsant, 1842
Sinodendron cylindricum aspromontanum G.Müller, 1938
DORCINIParry, 1864
BolbelasmusBoucomont,1911
DorcusMacLeay, 1819
[Bolbelasmus gallicus
*Dorcus musimon
*Bolbelasmus romanorum
Gené, 1836
*Dorcus parallelipipedus
*Bolbelasmus unicornis
Arnone & Massa, 2010
(Schrank, 1789)
(Linnaeus, 1758)
OdonteusSamouelle, 1819
Dorcus parallelepipedus truquii Mulsant, 1855
*Odonteus armiger
PLATYCERINI Mulsant, 1842
(Scopoli, 1772)
Odontaeus armiger (Scopoli, 1772)
PlatycerusGeoffroy, 1762
*Platycerus caprea
(Mulsant, 1842)]
Bolboceras armiger (Scopoli, 1772)
(Degeer, 1774)
*Platycerus caraboides
GEOTRUPIDAE Latreille, 1802
(Linnaeus, 1758)
GEOTRUPINAELatreille, 1802
LUCANINAELatreille, 1804
LUCANINI Latreille, 1804
ENOPLOTRUPINI Paulian, 1945
LucanusScopoli, 1763
Chelotrupes Jekel, 1966
*Chelotrupes hiostiusGenè, 1836
Subgen.Lucanus Scopoli, 1763
Typhoeus (Chelotrupes) hiostius Genè, 1836
Lucanus (Lucanus) cervus
(Linnaeus, 1758)
*Chelotrupes matutinalis(Baudi, 1870)
*Lucanus (Lucanus) cervus cervus(Linnaeus, 1758)
Typhoeus (Chelotrupes) hiostius var. matutinalis (Baudi, 1870)
Lucanus (Lucanus) tetraodon Thunberg, 1806
Typhaeus Leach, 1815
*Lucanus (Lucanus) tetraodon sicilianus Planet, 1899
*Typhaeus typhoeus
*Lucanus (Lucanus) tetraodon tetraodon Thunberg, 1806
TROGIDAEMacLeay, 1819
1 ( Linnaeus, 1758)
Typhoeus (s.str.) typhoeus (Linnaeus, 1758)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
Trypocopris (Trypocopris) pyrenaeus bartolii(G. Dellacasa,
GEOTRUPINI Latreille, 1802
1974)
AnoplotrupesJekel, 1866
*Anoplotrupes stercorosus
*Trypocopris (Trypocopris) pyrenaeus cyanicolor(Capra,
(Scriba, 1791)
1930)
*Trypocopris (Trypocopris) pyrenaeus splendens(Heer, 1841)
Baraudia López-Colón, 1995
*Baraudia geminata
Trypocopris (Trypocopris) vernalis
(Gené, 1839)
Thorectes geminatus (Gené, 1839)
1958)
Ceratophyus Fischer von Waldheim, 1824
*Ceratophyus rossii
(Linnaeus, 1758)
*Trypocopris (Trypocopris) vernalis apenninicus(Mariani,
*Trypocopris (Trypocopris) vernalis fauveli(Bedel, 1911)
Geotrupes (Trypocopris) vernalis obscurus Mulsant, 1842
Jekel, 1866
*Trypocopris (Trypocopris) vernalis vernalis(Linnaeus, 1758)
Ceratophyus fischeri Fischer von Waldheim ( sensu Boucomont, 1902)
Geotrupes Latreille, 1797
ZuninoeusLópez-Colón, 1989
Subgen.StereopygeA. Costa, 1847
Zuninoeus hoppei
*Geotrupes (Stereopyge) douei
(Sturm & Hagenbach, 1825)
Thorectes hoppei (Sturm & Hagenbach, 1825)
Gory, 1841
Stereopyge douei Gory, 1841
Subgen.GeotrupesLatreille, 1797
*Geotrupes (Geotrupes) mutator
Marsham, 1802
*Geotrupes (Geotrupes) spiniger
Marsham, 1802
HYBOSORIDAEErichson, 1847
HYBOSORINAE Erichson, 1847
Geotrupes puncticollis (Malinowsky, 1811)
*Geotrupes (Geotrupes) stercorarius
Hybosorus Macleay, 1819
(Linnaeus, 1758)
*Hybosorus illigeri
JekeliusLópez-Colón, 1989
Subgen. Jekelius
Reiche, 1853
Hybosorus arator Illiger,1803
López-Colón, 1989
*Jekelius (Jekelius) brullei brullei( Jekel, 1866)
OCHODAEIDAEMulsant & Rey, 1871
Thorectes brullei (Jekel, 1866)
*Jekelius (Jekelius) intermedius
( O.G. Costa, 1839)
OCHODAEINAEMulsant & Rey, 1871
Thorectes intermedius (O.G. Costa, 1839)
*Jekelius (Jekelius) sardous
(Erichson, 1847)
Ochodaeus Dejean, 1821
Thorectes sardous Erichson, 1847
*Ochodaeus chrysomeloides
Subgen. RudolfpetrovitziaRey & López-Colón, 2003
(Schrank, 1781)
Ochodaeus cychramoides Reitter, 1892
*Jekelius (Rudolfpetrovitzia) marginatus (Poiret, 1787)
Thorectes marginatus (Poiret, 1787)
Jekelius (Petrovitzia) marginatus (Poiret, 1787)
GLAPHYRIDAEMacLeay, 1819
SericotrupesZunino, 1984
Amphicoma Latreille, 1807
*Sericotrupes niger (Marsham, 1802)
Amphicoma abdominalis
Trypocopris Motschulsky, 1860
Subgen. Trypocopris
Motschulsky, 1860
Trypocopris (Trypocopris) alpinus
*Amphicoma abdominalis abdominalis(Fabricius, 1781)
(Sturm & Hagenbach, 1825)
Anthypna abdominalis abdominalis (Fabricius, 1781)
*Amphicoma abdominalis aemiliana(Ghidini, 1956)
*Trypocopris (Trypocopris) alpinus alpinus(Sturm &
Anthypna abdominalis aemiliana Ghidini, 1956
Hagenbach, 1825)
*Amphicoma abdominalis binaghii(Ghidini, 1956)
[Trypocopris (Trypocopris) alpinus balcanicolaMiksic, 1954]
Anthypna abdominalis binaghii Ghidini, 1956
*Trypocopris (Trypocopris) alpinus
*Amphicoma carcelii
marianiiF:\Scarabeoidea (Laporte de Castelnau, 1832)
Anthypna carceli Laporte de Castelnau, 1832
lug2014\scarabeidi\Trypocopris alpinus opacus.htm Loebl, 2006
Geotrupes (Trypocopris) alpinus opacus Mariani, 1958
Trypocopris (Trypocopris) pyrenaeus (Charpentier, 1825)
2 (Fabricius, 1781)
Anthypna abdominalis (Fabricius, 1781)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
AgoliusMulsant & Rey, 1870
SCARABAEIDAELatreille, 1802
Agolius abdominalis(Bonelli, 1812)
Aphodius mixtus Villa & Villa, 1833
CHIRONINAE
*Agolius abdominalis abdominalis(Bonelli, 1812)
C.E. Blanchard, 1845
Aphodius (Agolius) abdominalis abdominalis Bonelli, 1812
ChironMacleay, 1819
*Agolius abdominalis emilianus(G. Dellacasa, 1983)
[Chiron cylindrus (Fabricius, 1798)]
Aphodius (Agolius) abdominalis emilianus G. Dellacasa, 1983
Chiron digitatus (Fabricius, 1801)
AgrilinusMulsant & Rey, 1870
AEGIALIINAELaporte de Castelnau, 1840
*Agrilinus ater (Degeer, 1774)
Aphodius (Agrilinus) ater (Degeer, 1774)
AegialiaLatreille, 1807
*Agrilinus constans
[Aegialia arenaria (Fabricius, 1787)]
*Agrilinus convexus (Erichson, 1848)
PsammoporusThomson, 1859
*Psammoporus latipunctus
(Duftschmid, 1805)
Aphodius (Agrilinus) constans Duftschmidt, 1805
Aphodius (Agrilinus) ater convexus Erichson, 1848
*Agrilinus ibericus
Gredler, 1866
Psammoporus sabuleti auct. nec Panzer, 1797
(Harold, 1874)
Aphodius (Agrilinus) ibericus basilicatus Fiori, 1907
APHODIINAELeach, 1815
Alocoderus A. Schmidt, 1913
APHODIINILeach, 1815
*Alocoderus hydrochaeris
APHODIINALeach, 1815
Amidorus Mulsant & Rey, 1870
Acanthobodilus G. Dellacasa, 1983
*Acanthobodilus immundus
[Amidorus cribrarius
(Creutzer, 1799)
[Amidorus cribricollis
Acrossus Mulsant, 1842
*Amidorus immaturus
Aphodius (Acrossus) bimaculatus (Laxmann, 1770)
Amidorus obscurus
Aphodius (Amidorus) obscurus obscurus (Fabricius, 1792)
(Baudi, 1870)
*Amidorus obscurus latinus(G. Dellacasa, 1983)
Aphodius (Acrossus) laticollis Baudi, 1870
Aphodius (Amidorus) obscurus latinus G. Dellacasa, 1983
(Fabricius, 1775)
*Amidorus thermicola
Aphodius (Acrossus) luridus (Fabricius, 1775)
Aphodius (Acrossus) rufipes (Linnaeus, 1758)
Ammoecius Mulsant, 1842
(Harold, 1862)
*Ammoecius brevis
*Acrossus siculus siculus(Harold, 1862)
*Ammoecius elevatus
*Acrossus siculus buturensis(G. Dellacasa, 1983)
[Ammoecius numidicus
AgoliinusA. Schmidt, 1913
*Ammoecius rugifrons
(Erichson, 1848)]
Aubé, 1850
Anomius Mulsant & Rey, 1870
(Gyllenhal, 1808)
*Anomius castaneus(Illiger, 1803)
Aphodius (Agoliinus) piceus Gyllenhall, 1808
Aphodius (Anomius) castaneus Illiger, 1803
(Reitter, 1892)
*Anomius crovettii
Aphodius (Agoliinus) ragusae Reitter, 1892
(G. Dellacasa, 1983)
Aphodius (Anomius) crovettii G. Dellacasa, 1983
(Reitter, 1892)
AphodiusIlliger, 1798
Aphodius (Agoliinus) satyrus Reitter, 1892
3 Mulsant, 1851]
Aphodius (Ammoecius) rugifrons (Aubé, 1850)
Aphodius (Agrilinus) nemoralis (Erichson, 1848)
*Agoliinus satyrus
(Olivier, 1789)
Aphodius (Ammoecius) elevatus (Olivier, 1789)
Aphodius (Acrossus) siculus buturensis G. Dellacasa, 1983
*Agoliinus ragusae
Erichson, 1848
Aphodius (Ammoecius) brevis (Erichson, 1848)
Aphodius (Acrossus) siculus siculus Harold, 1863
*Agoliinus piceus
(Sturm, 1800)
Aphodius (Pseudacrossus) thermicola Sturm, 1800
(Linnaeus, 1758)
[Agoliinus nemoralis
(Fabricius, 1792)
*Amidorus obscurus obscurus(Fabricius, 1792)
Aphodius (Acrossus) depressus (Kugelann, 1792)
Acrossus siculus
(Mulsant, 1842)
Aphodius (Amidorus) immaturus Mulsant, 1842
*Acrossus depressus (Kugelann, 1792)
*Acrossus rufipes
(Lucas, 1846)]
Aphodius (Pseudacrossus) cribricollis Lucas, 1846
[Acrossus bimaculatus (Laxmann, 1770)]
*Acrossus luridus
(Brullé, 1832)]
Aphodius (Amidorus) cribrarius Brullé, 1832
= Aphodius (Acanthobodilus) immundus Creutzer, 1799
*Acrossus laticollis
(Fabricius, 1798)
Aphodius (Alocoderus) hydrochaeris (Fabricius, 1798)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Aphodius coniugatus
*Chilothorax distinctus
(Panzer, 1795)
Aphodius (Aphodius) conjugatus (Panzer, 1795)
*Aphodius fimetarius
Aphodius (Chilothorax) distinctus (O. F. Müller, 1776)
*Chilothorax lineolatus
(Linnaeus, 1758)
Aphodius (Aphodius) fimetarius (Linnaeus, 1758)
Aphodius pedellus Degeer, 1774
[Chilothorax melanostictus(W. Schmidt, 1840)]
Aphodius (Chilothorax) melanostictus W. Schmidt, 1840
Aphodius (Aphodius) foetidus (Herbst, 1783)
*Chilothorax paykulli
Aphodius scybalarius auct. nec Fabr. 1781
Aphodius (Chilothorax) paykulli Bedel, 1907
ColobopterusMulsant, 1842
Biralus Mulsant & Rey, 1870
*Biralus mahunkaorum (Adam, 1983)
*Colobopterus erraticus
(Herbst, 1789)
Coprimorphus Mulsant, 1842
Bodiloides M. Dellacasa & G. Dellacasa, 2005
*Coprimorphus scrutator
(Herbst, 1789)
Aphodius (Coprimorphus) scrutator (Herbst, 1789)
(Laicharting, 1781)
Aphodius nitidulus Fabricius, 1792
ErytusMulsant & Rey, 1870
*Bodiloides ictericus ghardimaouensis(Balthasar, 1929)
*Erytus aequalis
Aphodius (Bodilus) ictericus ghardimaouensis Balthasar, 1929
(A. Schmidt, 1907)
Aphodius (Erytus) aequalis A. Schmidt, 1907
Bodilus ictericus ghardimaouensis Balthasar, 1929
*Erytus cognatus
*Bodiloides ictericus ictericus(Laicharting, 1781)
(Fairmaire, 1860)
Aphodius (Erytus) klugi A. Schmidt, 1845
Aphodius (Bodilus) ictericus ictericus (Laicharting, 1781)
Bodilus ictericus ictericus (Laicharting, 1781)
EsymusMulsant & Rey, 1870
BodilopsisÁdám, 1994
*Esymus merdarius
(Fabricius, 1775)
Aphodius (Esymus) merdarius (Fabricius, 1775)
(Moll, 1782)
Esymus pusillus
Aphodius (Agrilinus) scybalarius (Fabricus, 1781)
Aphodius (Agrilinus) rufus (Moll, 1782)
(Herbst, 1789)
*Esymus pusillus pusillus(Herbst, 1789)
Agrilinus rufus (Moll, 1782)
Bodilopsis sordida
(Linnaeus, 1758)
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)
Aphodius (Biralus) satellitius (Herbst, 1789)
*Bodilopsis rufa
(Bedel, 1907)
Aphodius tessulatus Paykull, 1798
*Aphodius pedellus(De Geer, 1774)
Bodiloides ictericus
(Illiger, 1803)
Aphodius (Chilothorax) lineolatus Illiger, 1803
*Aphodius foetidus (Herbst, 1783)
*Biralus satellitius
(O.F. Muller, 1776)
Aphodius (Esymus) pusillus pusillus (Herbst, 1789)
(Fabricius, 1775)
Eudolus Mulsant & Rey, 1870
*Bodilopsis sordida sordida(Fabricius, 1775)
*Eudolus quadriguttatus
Aphodius (Agrilinus) sordidus (Fabricius, 1775)
(Herbst, 1783)
Aphodius (Eudolus) quadriguttatus (Herbst, 1783)
Agrilinus sordidus sordidus (Fabricius, 1775)
BodilusMulsant & Rey, 1870
EuheptaulacusG. Dellacasa, 1983
*Bodilus beduinus (Reitter, 1892)
Euheptaulacus carinatus
*Euheptaulacus carinatus carinatus(Germar, 1824)
Aphodius (Bodilus) beduinus Reitter, 1892
[Bodilus longispina
(Kuster, 1854)]
*Euheptaulacus carinatus brutius(Luigioni, 1920)
Aphodius (Bodilus) longispina Kuster, 1854
*Bodilus lugens
*Euheptaulacus carinatus esuriensG. Dellacasa, 1983
(Creutzer, 1799)
*Euheptaulacus porcellus
Aphodius (Bodilus) lugens (Creutzer, 1799)
[Euheptaulacus sus
CalamosternusMotschulsky, 1859
(Herbst, 1783)]
EuorodalusDellacasa, 1983
Aphodius (Calamosternus) hyxos algiricus Mariani & Pittino, 1983
*Euorodalus paracoenosus
(Linnaeus, 1767)
Aphodius (Calamosternus) granarius (Linnaeus, 1767)
(Balthasar & Hrubant, 1960)
Aphodius (Euorodalus) paracoenosus Balthasar & Hrubant, 1960
*Calamosternus mayeri (Pilleri, 1953)
*Euorodalus tersus
Aphodius (Calamosternus) mayeri Pilleri, 1953
(Erichson, 1848)
Aphodius (Euorodalus) tersus Erichson, 1848
Chilothorax Motschulsky, 1859
EupleurusMulsant, 1842
*Chilothorax conspurcatus
Eupleurus subterraneus
(Linnaeus, 1758)
Aphodius (Chilothorax) conspurcatus (Linnaeus, 1758)
4 (Frivaldsky, 1879)
*Euheptaulacus villosus (Gyllenhal, 1806)
*Calamosternus algiricus(Mariani & Pittino, 1983)
*Calamosternus granarius
(Germar, 1824)
(Linnaeus, 1758)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Eupleurus subterraneus subterraneus(Linnaeus, 1758)
*Melinopterus sphacelatus
Aphodius (Melinopterus) sphacelatus (Panzer, 1798)
Aphodius (Eupleurus) subterraneus subterraneus (Linnaeus, 1758)
*Melinopterus stolzi
HeptaulacusMulsant, 1842
*Heptaulacus rasettii
(Panzer, 1798)
(Reitter, 1906)
Aphodius (Melinopterus) stolzi Reitter, 1906
Carpaneto, 1978
*Heptaulacus testudinarius
*Melinopterus tingens
(Fabricius, 1775)
(Reitter, 1892)
Aphodius (Melinopterus) tingens Reitter, 1892
LabarrusMulsant & Rey, 1870
NeagoliusW. Koshantschikov, 1912
*Labarrus lividus
*Neagolius amblyodon
(Olivier, 1789)
Aphodius (Neagolius) amblyodon K. Daniel, 1900
Aphodius (Labarrus) lividus (Olivier, 1789)
*Neagolius bilimeki(Seidlitz, 1891)
LimarusMulsant & Rey, 1870
Aphodius (Neagolius) limbolarius limbolarius Reitter, 1892
[Limarus maculatus (Sturm, 1800)]
Aphodius (Neagolius) limbolarius danielorum A. Semenov, 1902
Neagolius limbolarius danielorum (A. Semenov, 1902)
Aphodius (Limarus) maculatus Sturm, 1800
*Limarus zenkeri
*Neagolius liguricus
(Germar, 1813)
*Neagolius montanus (Erichson, 1848)
1860
Aphodius (Neagolius) montanus Erichson, 1848
[Neagolius montivagus
*Liothorax niger(Illiger, 1798)
*Neagolius penninus (J. Daniel, 1902)
Aphodius (Liothorax) niger (Panzer, 1797)
Aphodius niger auctorum nec Panzer
Aphodius (Neagolius) penninus J. Daniel, 1902
Aphodius (Liothorax) wilsonae Maté & Angus, 2005
*Neagolius pollicatus
(Linnaeus, 1767)]
[Neagolius praecox (Erichson, 1848)]
LoraphodiusReitter, 1892
Aphodius (Neagolius) praecox Erichson, 1848
Neagolius schlumbergeri
(Faldermann, 1835)
Aphodius (Loraphodius) suarius Faldermann, 1835
(Seidlitz, 1891)
*Neagolius schlumbergeri consobrinus(K. Daniel, 1900)
Aphodius (Neagolius) schlumbergeri consobrinus K. Daniel, 1900
LoraspisMulsant & Rey, 1870
[Loraspis frater
(Erichson, 1848)
Aphodius (Neagolius) pollicatus Erichson, 1848
Aphodius (Liothorax) plagiatus (Linnaeus, 1767)
*Loraphodius suarius
(Erichson, 1848)]
Aphodius (Neagolius) montivagus Erichson, 1848
Liothorax muscorum (Ádám, 1994)
[Liothorax plagiatus
(J. Daniel, 1902)
Aphodius (Neagolius) liguricus J. Daniel, 1902
Aphodius (Limarus) zenkeri Germar, 1813
LiothoraxMotschulsky,
(K. Daniel, 1900)
*Neagolius schlumbergeri samniticus(J. Daniel, 1902)
(Mulsant & Rey, 1870)]
Aphodius (Neagolius) schlumbergeri samniticus J. Daniel, 1902
Aphodius (Loraspis) frater Mulsant & Rey, 1870
NialusMulsant & Rey, 1870
Mecynodes Mulsant & Rey, 1870
*Nialus varians
*Mecynodes striatulus(Waltl, 1835)
(Duftschmid, 1805)
Aphodius (Nialus) varians Duftschmid, 1805
Aphodius (Mecynodes) striatulus Waltl, 1835
= Aphodius parisii Schatzmayr, 1937
Nimbus Mulsant & Rey, 1870
Megatelus Reitter, 1892
[Nimbus affinis
(Panzer, 1823)]
Aphodius (Nimbus) affinis Panzer, 1823
*Megatelus contractus(Klug, 1845)
Aphodius (Nimbus) hoberlandti Tesar, 1945
Aphodius (Megatelus) contractus scolytoides Lucas, 1846
*Nimbus contaminatus
Megatelus contractus scolytoides (Lucas, 1846)
(Herbst, 1783)
Aphodius (Nimbus) contaminatus (Herbst, 1783)
Melinopterus Mulsant, 1842
*Melinopterus consputus
*Nimbus franzinii (Pittino, 1978)
Aphodius (Nimbus) franzinii Pittino, 1978
(Creutzer, 1799)
*Nimbus johnsoni (Baraud, 1976)
Aphodius (Melinopterus) consputus (Creutzer, 1799)
*Melinopterus prodromus
Aphodius (Nimbus) johnsoni Baraud, 1976
(Brahm, 1790)
*Nimbus obliteratus
Aphodius (Melinopterus) prodromus (Brahm, 1790)
*Melinopterus punctatosulcatus punctatosulcatus (Sturm,
[Nimbus proximus Ádám, 1994]
1805)
Aphodius (Melinopterus) punctatosulcatus Sturm, 1805
NobiusMulsant & Rey, 1870
Aphodius (Melinopterus) sabulicola Thomson, 1868
*Melinopterus reyi
*Nobius serotinus
(Reitter, 1892)
Aphodius (Melinopterus) reyi Reitter, 1892
(Panzer, 1799)
Aphodius (Nobius) serotinus (Panzer, 1799)
5 (Panzer, 1823)
Aphodius (Nimbus) obliteratus Panzer, 1823
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
Aphodius (Aphodius) aestivalis Stephens, 1839
OromusMulsant & Rey, 1870
Aphodius aestivalis Stephens, 1839
*Oromus alpinus
(Scopoli, 1763)
Aphodius foetens (Fabricius, 1787)
Aphodius (Oromus) alpinus (Scopoli, 1763)
SigorusMulsant & Rey, 1870
Otophorus Mulsant, 1842
*Sigorus porcus
*Otophorus haemorrhoidalis
(Linnaeus, 1758)
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)
SubrinusMulsant & Rey, 1870
Oxyomus Dejean, 1833
*Subrinus sturmi
*Oxyomorus sylvestris(Scopli, 1763)
Aphodius (Subrinus) sturmi Harold, 1870
TeuchestesMulsant, 1842
Aphodius (Parammoecius) corvinus Erichson, 1848
(Germar in Ahrens, 1817)
*Teuchestes fossor
Aphodius (Parammoecius) gibbus Germar, 1817
TrichonotulusBedel, 1911
Aphodius (Parammoecius) pyrenaeus J. du Val, 1859
*Trichonotulus scrofa
PhalacronothusMotschulsky, 1859
VolinusMulsant & Rey, 1870
Aphodius (Phalacronothus) biguttatus Germar, 1824
*Volinus sticticus (Panzer, 1798)
(Linnaeus, 1761)
*Phalacronothus quadrimaculatus
Aphodius equestris Panzer, 1798
quadrimaculatus(Linnaeus, 1761)
Aphodius (Chilothorax) sticticus (Panzer, 1798)
Aphodius (Phalacronothus) quadrimaculatus quadrimaculatus (Linnaeus,
[EUPARIINI A. Schmidt, 1910]
1761)
Ataenius Harold, 186
PlagiogonusMulsant, 1842
[Ataenius picinusHarold, 1867]
(Olivier, 1789)
PSAMMODIINIMulsant, 1840
Aphodius (Plagiogonus) putridus Geoffroy in Fourcroy, 1785
PSAMMODIINAMulsant, 1840
PlanolinoidesM. Dellacasa & G. Dellacasa, 2005
*Planolinoides borealis
(Fabricius, 1787)
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)
*Phalacronothus biguttatus(Germar, 1824)
*Plagiogonus arenarius
(Linnaeus, 1758)
= Aphodius (Teuchestes) fossor (Linnaeus, 1758)
(J. du Val, 1859)
Phalacronothus quadrimaculatus
(Klug, 1845)
Aphodius (Subrinus) vitellinus Klug, 1845
*Parammoecius corvinus (Erichson, 1848)
*Parammoecius pyrenaeus
(Harold, 1870)
*Subrinus vitellinus
Parammoecius Seidlitz, 1891
*Parammoecius gibbus
(Fabricius, 1792)
Aphodius (Sigorus) porcus (Fabricius, 1792)
BrindalusLandin, 1960
(Gyllenhal, 1827)
*Brindalus porcicollis
Aphodius (Planolinus) borealis Gyllenhal, 1827
Planolinus borealis (Gyllenhal, 1827)
(Illiger, 1803)
*Brindalus rotundipennis
PlanolinusMulsant & Rey, 1870
DiastictusMulsant, 1842
*Planolinus fasciatus
*Diastictus vulneratus
(Olivier, 1789)
(Reitter, 1892)
(Sturm, 1805)
Aphodius (Planolinus) putridus (Herbst, 1789)
PsammodiusFallén, 1807
Aphodius (Planolinus) uliginosus (Hardy, 1847)
Aphodius (Planolinus) tenellus sensu auctorum nec Say, 1823
*Psammodius asper
PseudacrossusReitter, 1892
(Fabricius, 1775)
Psammodius sulcicollis Illiger, 1802
*Psammodius basalis
*Pseudacrossus suffertus(A. Schmidt, 1922)
(Mulsant & Rey, 1870)
Aphodius (Pseudacrossus) suffertus ampliatus Reitter, 1892
*Psammodius laevipennis
Aphodius (Pseudacrossus) suffertus defiorei Fiori, 1907
*Psammodius nocturnus Reitter, 1892
PubinusMulsant & Rey, 1870
*Psammodius pierottii
[Pubinus tomentosus(O.F.Muller, 1776)]
*Psammodius plicicollis
Aphodius (Pubinus) tomentosus (O.F.Muller, 1776)
Erichson, 1848
PlatytomusMulsant, 1842
(Fabricius, 1787)
*Platytomus laevistriatus
Aphodius (Aphodius) foetens (Fabricius, 1787)
6 Pittino, 1979
RHYSSEMINAPittino & Mariani, 1986
RhodaphodiusÁdám, 1994
*Rhodaphodius foetens
A. Costa, 1844
(Perris, 1869)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Platytomus tibialis
*Gymnopleurus mopsus mopsus
(Fabricius, 1798)
*Gymnopleurus sturmii
PleurophorusMulsant, 1842
*Pleurophorus caesus
SYSIPHINIMulsant, 1842
*Pleurophorus maghrebinicus Pittino & Mariani, 1986
SisyphusLatreille, 1807
*Pleurophorus mediterranicus Pittino & Mariani, 1986
Sisyphus schaefferi
Reitter, 1893
*Pleurophorus pannonicus
COPRINILeach, 1815
*Rhyssemus annaedicatus
CoprisGeoffroy, 1762
Pierotti, 1980
*Copris lunaris
(Linnaeus, 1767)
*Copris hispanus cavolinii(Petagna, 1792)
Rhyssemus algiricus marqueti Reiche, 1863
*Rhyssemus plicatus
*Copris umbilicatus
Reitter, 1893
(Germar, 1817)
BubasDejean, 1833
(Olivier, 1789)
*Bubas bison
Rhyssemus godarti Mulsant & Rey, 1870
*Rhyssemus verrucosus
Mulsant, 1842
[Trichiorhyssemus setulosus
(Linnaeus, 1767)
[Bubas bubaloidesJanssens, 1938]
TrichiorhyssemusClouet, 1901
*Trichiorhyssemus dalmatinus
Abeille de Perrin, 1901
ONITINILaporte de Castelnau, 1840
*Rhyssemus sardous Pierotti, 1980
*Rhyssemus sulcatus
(Linnaeus, 1764)
*Copris hispanus hispanus(Linnaeus, 1764)
Reiche, 1863
*Rhyssemus parallelus
(Linnaeus, 1758)
Copris hispanus
*Rhyssemus limbolarius Petrovitz, 1963
*Rhyssemus marqueti
(Linnaeus, 1758)
*Sisyphus schaefferi schaefferi(Linnaeus, 1758)
Petrovitz, 1961
RhyssemusMulsant, 1842
*Rhyssemus germanus
MacLeay, 1821
Gymnopleurus sturmi MacLeay, 1821
Creutzer in Panzer, 1796)
*Pleurophorus opacus
(Pallas, 1781)
CheironitisLansberge, 1875
Petrovitz, 1967
*Cheironitis furcifer
Reitter, 1892)]
(Rossi, 1792)
Chironitis furcifer (Rossi, 1792)
*Cheironitis irroratus
SCARABAEINAELatreille, 1802
(Rossi, 1790)
Chironitis irroratus (Rossi, 1790)
[Cheironitis ungaricus
SCARABAEINILatreille, 1802
(Herbst, 1789)]
Chironitis hungaricus (Herbst, 1789)
ScarabaeusLinnaeus, 1758
OnitisFabricius, 1798
Subgen. Ateuchetus Bedel, 1892
Onitis alexis
*Scarabaeus(Ateuchetus) laticollis(Linnaeus, 1767)
Klug, 1835
*Onitis alexis septentrionalisBalthasar, 1942
Ateuchetus laticollis Linnaeus, 1767
*Scarabaeus (Ateuchetus) semipunctatus(Fabricius, 1792)
*Onitis belial Fabricius, 1798
Ateuchetus semipunctatus Fabricius, 1792
*Onitis ion
(Olivier, 1789)
*Scarabaeus (Ateuchetus) variolosus(Fabricius, 1787)
ONITICELLINIKolbe, 1905
Ateuchetus variolosus Fabricius, 1787
Subgen. Scarabaeus Linnaeus, 1758
EuoniticellusJanssens, 1953
*Scarabaeus (Scarabaeus) pius(Illiger, 1803)
* Scarabaeus (Scarabaeus) sacer
*Euoniticellus fulvus (Goeze, 1777)
Linnaeus, 1758
[Euoniticellus intermedius
*Scarabaeus (Scarabaeus) typhon(Fischer von Waldheim, 1823)
Scarabaeus affinis Brullé, 1832
*Euoniticellus pallens
GYMNOPLEURINI Lacordaire, 1856
ONTHOPHAGINIBurmeister, 1846
(Fabricius, 1787)
CaccobiusThomson, 1859
Gymnopleurus coriarius (Herbst, 1789)
Gymnopleurus geoffroyi
*Caccobius schreberi
(Fuesslin, 1775)
EuonthophagusBalthasar, 1959
(Pallas, 1781)
7 (Linnaeus, 1767)
Histeridium schreberi (Linné, 1767)
*Gymnopleurus geoffroyi geoffroyi(Fuesslin, 1775)
Gymnopleurus mopsus
(Olivier, 1789)
*Euoniticellus pallipes (Fabricius, 1781)
GymnopleurusIlliger, 1803
*Gymnopleurus flagellatus
(Reiche, 1849)]
Oniticellus speciosus A. Costa, 1853
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
Euonthophagus amyntas
Chaetonyx robustus Schaum, 1862
(Olivier, 1789)
*Euonthophagus amyntas amyntas(Olivier, 1789)
*Chaetonyx robustus italicusMariani, 1946
[Euonthophagus crocatus crocatus(Mulsant & Godart, 1873)]
*Chaetonyx robustus liguricusMariani, 1946
*Euonthophagus gibbosus
(Scriba, 1790)
HybalusDejean, 1833
OnthophagusLatreille, 1802
*Hybalus benoitiTournier, 1864
Subgen. Furconthophagus Zunino, 1979
*Hybalus bigibber
*Onthophagus (Furconthophagus) furcatus
[Hybalus cornifrons Brullé, 1832]
( Fabricius, 1781)
Hybalus graecus Sturm, 1843
Subgen. Onthophagus Latreille, 1802
*Onthophagus (Onthophagus) illyricus (Scopoli, 1763)
*Onthophagus (Onthophagus) taurus
MELOLONTHINAE
(Schreber, 1759)
*Onthophagus (Palaeonthophagus) andalusicus
HymenopliaEschscholtz, 1830
Waltl, 1835
*Hymenoplia sicula
Onthophagus andalusicus italicus Goidanich, 1925
*Onthophagus (Palaeonthophagus) baraudiNicolas, 1964
Subgen. Maladera Mulsant & Rey, 1871
(Preyssler,
*Maladera (Maladera) holosericea
1790)
Onthophagus (Palaeonthophagus) gibbulus
(Pallas, 1781)
Subgen. Omaloplia
(Pallas, 1781)
*Onthophagus (Palaeonthophagus) grossepunctatus
(Scopoli, 1772)
Omaloplia Schonherr, 1817
*Onthophagus (Palaeonthophagus) gibbulus gibbulus
Stephens, 1830
[Omaloplia (Omaloplia) hericia
Reitter,
Chobaut, 1907]
Homaloplia (Homaloplia) hericius Chobaut, 1907
1905
*Onthophagus (Palaeonthophagus) joannae
*Omaloplia (Omaloplia) lonae
Goljan, 1953
*Onthophagus (Palaeonthophagus) lemur
(Fabricius, 1781)
*Onthophagus (Palaeonthophagus) massai
Baraud, 1975
*Onthophagus (Palaeonthophagus) nuchicornis
Schatzmayr, 1923
Homaloplia (Homaloplia) lonae Schatzmayr, 1923
Omaloplia (Omaloplia) ruricola
Fabricius, 1775
Homaloplia (Homaloplia) ruricola (Fabricius,1775)
*Onthophagus (Palaeonthophagus) medius(Kugelann, 1792)
*Omaloplia (Omaloplia) ruricola nicolasiBaraud, 1965
(Linnaeus,
Homaloplia (Homaloplia) nicolasi Baraud, 1965
*Omaloplia (Omaloplia) ruricola ruricolaFabricius, 1775
1758)
*Onthophagus (Palaeonthophagus) opacicollis
Reitter, 1892
Homaloplia (Homaloplia) nicolasi tergestina Baraud, 1965
Onthophagus schatzmayri Pierotti, 1959
Paratriodonta Baraud, 1962
*Onthophagus (Palaeonthophagus) ovatus (Linnaeus, 1767)
*Onthophagus (Palaeonthophagus) ruficapillus
Brullé, 1832
*Onthophagus (Palaeonthophagus) semicornis
(Panzer, 1798)
*Onthophagus (Palaeonthophagus) vacca
*Paratriodonta cinctipennis
*Paratriodonta romana
*Onthophagus (Palaeonthophagus) verticicornis
(Lucas, 1846)
(Brenske, 1890)
SericaMacleay, 1819
(Linnaeus, 1767)
Subgen. Serica Macleay, 1819
(Laicharting,
1781)
*Serica (Serica) brunnea (Linnaeus, 1758)
Onthophagus nutans Fabricius, 1787
[Onthophagus (Palaeonthophagus) vitulus
Serica brunnea var. Berardii Luigioni, 1928
(Fabricius, 1776)]
TriodontellaReitter, 1919
Zunino, 1979
[Onthophagus (Parentius) punctatus
*Triodontella alni
(Illiger, 1803)]
Subgen. Trichonthophagus
(Blanchard, 1850)
Triodonta alni Blanchard, 1850
Onthophagus (Relictonthophagus) punctatus (Illiger, 1803)
Triodontella alni luteipes (Fairmaire, 1881)
Zunino, 1979
*Triodontella bucculenta(Baraud, 1962)
*Onthophagus (Trichonthophagus) maki (Illiger, 1803)
Triodonta bucculenta Baraud, 1962,
*Triodontella mimulaLeo & Fancello, 2007
ORPHNINAEErichson, 1847
*Triodontella nitidula
ORPHNINIErichson, 1847
(Rossi, 1790)
Triodonta nitidula (Rossi, 1790)
Chaetonyx Schaum, 1862
*Triodontella raymondi
8 Blanchard, 1850
MaladeraMulsant & Rey, 1871
*Onthophagus (Palaeonthophagus) coenobita(Herbst, 1783)
*Onthophagus (Palaeonthophagus) fracticornis
Leach in Samouelle, 1819
SERICINIKirby, 1837
Subgen. Palaeonthophagus Zunino, 1979
Subgen. Parentius
Reitter, 1892
(Perris, 1869)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
Triodonta raymondi Perris, 1869
*Triodontella sardoa
Amphimallon solstitiale javeti Stierlin, 1864
Amphimallon majale
(Baraud, 1962)
Triodonta sardoa Baraud, 1962
HOPLIINILatreille, 1829
Subgen. Decamera
Zur Strassen, 1954]
*Amphimallon ochraceum
(Knoch, 1801)
(Olivier, 1789)
*Amphimallon pseudomajale
Mulsant, 1842
*Hoplia (Decamera) fiorii
Amphimallon solstitiale
Fracassi, 1906
Hoplia (Decamera) philanthus
(Fuesslin, 1775)
*Hoplia (Decamera) philanthus philanthus
*Hoplia (Decamera) praticola
*Amphimallon vitalei
Duftschmid, 1805
[Aplidia attenuata (Reiche, 1852]
*Aplidia etrusca(Kraatz, 1882)
(Poda, 1761)
Hoplia farinosa (Linnaeus, 1761)
Haplidia etrusca Kraatz, 1882
*Aplidia hirticollis(Burmeister, 1855)
Massa, 1979
*Hoplia (Hoplia) brunnipes
Haplidia hirticollis Burmeister, 1855
Bonelli, 1812
*Aplidia massai(Baraud, 1975)
(Rossi, 1792)
[Hoplia (Hoplia) flavipes
Haplidia massai Baraud, 1975
Germar, 1824]
*Hoplia (Hoplia) graminicola
Aplidia transversa (Fabricius, 1801)
(Fabricius, 1792)
*Hoplia (Hoplia) maremmana
*Hoplia (Hoplia) minuta
Luigioni, 1932
Haplidia attenuata Reiche, 1852)
*Hoplia (Hoplia) argentea
*Hoplia (Hoplia) dubia
(Linnaeus, 1758)
AplidiaHope, 1837
(Fuesslin, 1775)
Illiger, 1803
*Hoplia (Hoplia) attilioi
Sabatinelli, 1977
*Amphimallon solstitiale solstitiale (Linnaeus, 1758)
Hoplia farinosa auctorum nec Linnaeus 761
Subgen. Hoplia
[Amphimallon montanum
*Amphimallon pini
HopliaIlliger, 1803
(Razoumowsky, 1789)
Haplidia transversa (Fabricius, 1801)
Leo, Liberto, Rattu & Sechi, 2010
*Aplidia transversa transversa(Fabricius, 1801)
Panzer, 1789
*Hoplia (Hoplia) paganettii
J. Muller, 1907
*Hoplia (Hoplia) pubicollis
Küster, 1849
*Hoplia (Hoplia) walterrossii
*Aplidia villigera (Burmeister, 1855)
Haplidia villigera Burmeister, 1855
FirminusCoca-Abia, 2003
Sabatinelli in Baraud, 1992
*Firminus baudii
(Brenske, 1886)
Rhizotrogus baudii Brenske, 1886
*Firminus bellieri (Reiche, 1862)
RHIZOTROGINIBurmeister, 1855
Rhizotrogus bellieri Reiche, 1862
AmadotrogusReitter, 1902
*Amadotrogus grassii
Firminus ciliatus
(Mainardi, 1902)
*Firminus ciliatus ciliatus (Reiche, 1862)
Rhizotrogus grassii Mainardi, 1902
*Amadotrogus insubricus
*Firminus ciliatus vexillis(Reitter, 1902)
(Burmeister, 1855)
Rhizotrogus ciliatus vexillis Reitter, 1902
Rhizotrogus insubricus Burmeister, 1855
*Firminus fossulatus (Mulsant & Rey, 1859)
Rhizotrogus fiorii Brenske, 1893
*Amadotrogus quercanus (Burmeister, 1855)
Rhizotrogus fossulatus Mulsant & Rey, 1859
*Firminus procerus
Amphimallon (Amadotrogus) quercanum Burmeister, 1855
*Amadotrogus tarsalis
(Reiche, 1862)
Geotrogus procerus (Baudi, 1870)
(Mulsant, 1842)
GeotrogusGuérin-Menéville, 1842
Rhizotrogus vicinus Mulsant, 1842
*Geotrogus euphytus lamantiai
Rhizotrogus rugifrons Burmeister, 1855
*Amphimallon atrum
Pseudoapterogyna euphytus lamantiai Sparacio, 2014
Rhizotrogus insularis Reiche, 1862
Brenske, 1886
*Geotrogus michaelis (Sparacio, 2014)
(Scopoli, 1786)
*Amphimallon gianfranceschii
Pseudoapterogyna michaelis Sparacio, 2014
*Geotrogus pellegrinensis
Luigioni, 1931
(Brenske in Ragusa, 1893)
Pseudoapterogyna pellegrinensis (Brenske in Ragusa, 1893)
Stierlin, 1864
9 (Blanchard, 1851)
Rhizotrogus genei Blanchard, 1851
(Herbst, 1790)
*Amphimallon fuscum
*Amphimallon javeti
*Geotrogus genei
(Herbst, 1790)
*Amphimallon burmeisteri
(Sparacio, 2014)
Pseudoapterogyna euphytus (Buquet, 1840)
AmphimallonLatreille, 1825
*Amphimallon assimile
(Baudi, 1870)
Rhizotrogus procerus Baudi, 1870
Rhizotrogus tarsalis Reiche, 1862
*Amadotrogus vicinus
(Reiche, 1862)
Rhizotrogus ciliatus Reiche, 1862
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Geotrogus sicelis
Blanchard, 1851
*Anoxia (Protanoxia) orientalis
*Geotrogus vorax
(Marseul, 1878)
MelolonthaFabricius, 1775
Pseudoapterogyna vorax (Marseul, 1878)
*Melolontha hippocastani
HolochelusReitter, 1889
Subgen.
*Melolontha melolontha
Miltotrogus Reitter, 1902
*Holochelus (Miltotrogus) fraxinicola
*Melolontha pectoralis
(Sturm & Hagenbach,
Miltotrogus fraxinicola (Sturm & Hagenbach, 1825)
*Holochelus (Miltotrogus) vernus
Subgen. Polyphylla
Harris, 1841
*Rhizotrogus cicatricosus
(Linnaeus, 1758)
Polyphylla (Polyphylla) ragusae
*Rhizotrogus aestivus (Olivier, 1789)
Kraatz, 1882
Polyphylla ragusai auctt.
Mulsant, 1842
*Polyphylla (Polyphylla) ragusae ragusaeKraatz, 1882
Rhizotrogus praecox Leoni, 1906
*Polyphylla (Polyphylla) ragusae aliquoiMassa & Tassi in
*Rhizotrogus maculicollisF:\Scarabeoidea Baraud, 1977
lug2014\scarabeidi\Rhizotrogus TANYPROCTINI Erichson, 1847
(= PACHYDEMINIBurmeister, 1855)
maculicollis.htm (A. & G.B. Villa, 1833)
ElaphoceraGené, 1836
Mulsant, 1842
*Rhizotrogus pallidipennis
*Elaphocera emarginata (Gyllenhal, 1817)
Blanchard, 1851
*Elaphocera erichsoni
Sabatinelli, 1975
*Rhizotrogus sassariensis
Drumont, Muret, Hayer & Penner, 1999
*Polyphylla (Polyphylla) fullo
RhizotrogusLatreille, 1825
*Rhizotrogus romanoi
Megerle von Muhlfeld, 1812
PolyphyllaHarris, 1841
(Germar, 1824)
Miltotrogus vernus (Germar, 1824)
*Rhizotrogus marginipes
Fabricius, 1801
(Linnaeus, 1758)
*Melolontha sardiniensis
1825)
(Krynicki, 1832)
Harold in Gemminger & Harold, 1869
Elaphocera erichsoni J. du Val, 1860
Perris, 1869
LiogenysGuèrin-Méneville, 1830
*Rhizotrogus siculusF:\Scarabeoidea Peritryssus Reitter, 1918
lug2014\scarabeidi\Rhizotrogus siculus.htm Baraud,
[Liogenys excisa
1970
(Reitter, 1918)]
Peritryssus excisus Reitter, 1918
MELOLONTHINILeach in Samouelle, 1819
PachydemaLaporte de Castelnau, 1832
AnoxiaLaporte de Castelnau, 1832
*Pachydema hirticollis
Subgen. Anoxia Laporte de Castelnau, 1832
*Anoxia (Anoxia) pilosa
RUTELINAE
(Fabricius, 1792)
Anoxia (Anoxia) scutellaris
ANOMALINA Streubel, 1839
*Anoxia (Anoxia) scutellaris argenteaAliquò & Massa, 1976
AnomalaSamouelle, 1819
*Anoxia (Anoxia) scutellaris siculaMotschulsky, 1860
*Anomala ausonia Erichson, 1847
(Fabricius, 1781)
*Anomala devota
*Anoxia (Anoxia) villosa villosa(Fabricius, 1781)
*Anomala dubia
( Schönherr, 1817)
Anoxia (Mesanoxia) matutinalis
*Anomala vitis
(Fabricius, 1775)
Exomala Reitter, 1903
*Anoxia (Mesanoxia) matutinalis matutinalisLaporte de
Subgen. Neoblitopertha
*Anoxia (Mesanoxia) matutinalis corsicanaSabatinelli, 1976
Baraud, 1991
*Exomala (Neoblitopertha) succincta(Laporte de Castelnau, 1840)
*Anoxia (Mesanoxia) matutinalis moltoniiSabatinelli, 1976
Exomala (Neoblitopertha) campestris (Latreille, 1804)
*Exomala (Neoblitopertha) leonii
*Anoxia (Mesanoxia) matutinalis sardoaMotschulsky, 1860
Blitopertha leonii Luigioni, 1933
*Anoxia (Mesanoxia) matutinalis thumseri Guerlach, 2011
MimelaKirby, 1823
Medvedev, 1951
10 (Scopoli, 1763)
Laporte de Castelnau, 1832
Castelnau, 1832
Subgen. Protanoxia
(Rossi, 1790)
Anomala (Dichomala) devota (Rossi, 1790)
Subgen. Mesanoxia Medvedev, 1951
*Anoxia (Mesanoxia) australis
MacLeay, 1819
ANOMALINIStreubel, 1839
Mulsant, 1842
*Anoxia (Anoxia) scutellaris scutellarisMulsant, 1842
Anoxia (Anoxia) villosa
(Fabricius, 1787)
(Luigioni, 1933)
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Mimela aurata
Mimela junii
*Calicnemis latreillii
(Fabricius, 1801)
Callicnemis latreillei Laporte de Castelnau, 1832
(Duftschmid, 1805)
*Calicnemis sardiniensis
*Mimela junii junii(Duftschmid, 1805)
PentodonHope, 1837
Mimela junii calabrica Machatschke 1952
*Mimela junii gigliocolaMachatschke, 1957
Pentodon algerinus
Mimela junii giglicola Machatschke, 1952
(Fuesslin, 1778)
Pentodon algerinum (Herbst, 1789)
*Mimela junii miksiciSparacio, 2003
*Pentodon algerinus algerinus(Fuesslin, 1778)
*Mimela junii rugosulaFairmaire, 1859
Pentodon bispinosum Küster, 1852
Pentodon bidens
Mimela junii corsicana Machatschke, 1957
Mimela junii corsicana Machatschke, 1952
(Pallas, 1771)
*Pentodon bidens punctatus(Villers, 1789)
PhylloperthaStephens, 1830
Pentodon punctatum (Villers, 1789)
Phyllognathus Eschscholtz, 1830
(Linnaeus, 1758)
*Phyllognathus excavatus
ANISOPLIINA Burmeister, 1844
ORYCTINIBurmeister, 1847
Schönherr, 1817
*Anisoplia (Anisoplia) agricola(Poda, 1761)
OryctesIlliger, 1798
*Anisoplia (Anisoplia) bromicola
Subgen. Oryctes Illiger, 1798
(Germar, 1817)
Oryctes (Oryctes) nasicornis (Linnaeus, 1758)
*Anisoplia (Anisoplia) erichsoniReitter, 1889
*Oryctes (Oryctes) nasicornis corniculatusVilla & Villa, 1833
[Anisoplia (Anisoplia) flavipennisBrullé, 1832]
Anisoplia (Anisoplia) monticola
Oryctes nasicornis laevigatus Heer, 1841
Erichson, 1847
*Oryctes (Oryctes) nasicornis grypusIlliger, 1803
*Anisoplia (Anisoplia) monticola monticola Erichson, 1847
Anisoplia neapolitana Reitter, 1889
CETONIINAELeach, 1815
*Anisoplia (Anisoplia) monticola marginataKraatz, 1883
*Anisoplia (Anisoplia) sabatinellii
Baraud, 1991
*Anisoplia (Anisoplia) tempestiva
Erichson, 1847
*Anisoplia (Anisoplia) villosa
VALGINIMulsant, 1842
ValgusScriba, 1790
*Valgus hemipterus
(Goeze, 1777)
Anisoplia simplicifronsReitter, 1889
[Subgen. Autanisoplia
TRICHIINAFleming, 1821
(Herbst, 1783)]
[Anisoplia (Autanisoplia) austriaca austriaca(Herbst, 1783)]
GnorimusLepeletier & Serville, 1828
HemichaetopliaBaraud, 1986
*Hemichaetoplia pallidipennis
*Gnorimus decempunctatus
(Gyllenhal, 1817)
*Gnorimus nobilis nobilis(Linnaeus, 1758)
Aleurostictus nobilis (Linnaeus, 1758)
*Gnorimus variabilis
Erichson, 1840
(Linnaeus, 1758)
Gnorimus octopunctatus (Fabricius, 1775)
Aleurostictus variabilis (Linnaeus, 1758)
Dejean, 1821
*Pachypus caesus
Helfer, 1833
Aleurostictus decempunctatus (Helfer, 1833)
Anisoplia pallidipennis hypocrita(Burmeister, 1844)
PACHYPODINAE
(Linnaeus, 1758)
TRICHIINIFleming, 1821
Medvedev, 1949]
[Anisoplia (Autanisoplia) austriaca
Pachypus
(Forster, 1771)
Phyllognathus silenus (Fabricius, 1775)
Anisoplia Schönherr, 1817
Subgen. Anisoplia
Leo, 1985
Callicnemis sardiniensis Leo, 1985
*Mimela junii calabricaMachatschke, 1957
*Phyllopertha horticola
Laporte de Castelnau, 1832
TrichiusFabricius, 1775
Erichson, 1840
*Pachypus candidae (Petagna, 1787)
*Trichius fasciatus (Linnaeus, 1758)
*Pachypus melonii
*Trichius gallicus gallicus Dejean, 1821
Sparacio, 2008
*Pachypus sardiniensis
DYNASTINAE
Trichius rosaceus rosaceus (Voet, 1766)
Guerlach, Bazzato & Cillo, 2013
*Trichius gallicus zonatus
MacLeay, 1819
PENTODONTINI Mulsant, 1842
*Trichius sexualis
CalicnemisLaporte de Castelnau, 1832
11 Germar, 1831
Trichius rosaceus zonatus Germar, 1831
Bedel, 1906
Antonio Rey & Alberto Ballerio. Checklist degli Scarabaeoidea Italiani. Luglio 2014 ©
*Protaetia (Potosia) angustata angustata(Germar, 1817)
OSMODERMATINA Schenkling, 1922
Protaetia (Potosia) cuprea (Fabricius, 1775)
OsmodermaLepeletier & Serville, 1828
*Osmoderma cristinae
*Protaetia (Potosia) cuprea cuprea(Fabricius, 1775)
Sparacio, 1994
*Protaetia (Potosia) cuprea hypocrita(Ragusa, 1905)
Osmoderma eremita meridionale Tauzin, 1994
*Osmoderma eremita
Protaetia (Potosia) cuprea incerta (A. Costa, 1852)
(Scopoli, 1763)
*Protaetia (Potosia) cuprea metallica(Herbst, 1782)
Osmoderma eremitum (Scopoli, 1763)
*Osmoderma italicum
*Protaetia (Potosia) cuprea obscura(Andersch, 1797)
Sparacio, 2000
Protaetia (Potosia) fieberi
Osmoderma italicaSparacio, 2000
*Protaetia (Potosia) fieberi fieberi(Kraatz, 1880)
CETONIINILeach, 1815
*Protaetia (Potosia) opaca
CETONIINALeach, 1815
Subgen. Epicometis Burmeister, 1842
(Fabricius, 1787)
Tropinota (Epicometis) hirta (Poda, 1761)
Aethiessa squamosa (Gory & Percheron, 1833)
*Tropinota (Epicometis) hirta hirta(Poda, 1761)
CetoniaFabricius, 1775
Cetonia aurata
Epicometis hirta hirta (Poda, 1761)
*Tropinota (Epicometis) paulae Leo, 2010
(Linné, 1758)
*Cetonia aurata aurata (Linnaeus, 1758)
Subgen. Tropinota Mulsant 1842
*Cetonia aurata pisanaHeer, 1841
Tropinota (Tropinota) squallida
*Cetonia aurata sicula
Cetonia carthami
*Tropinota (Tropinota) squallida squallida
Gory & Percheron, 1833
Tropinota squalida (Scopoli, 1763)
LEUCOCELINAKraatz, 1882
ProtaetiaBurmeister, 1842
OxythyreaMulsant, 1842
Subgen. Potosia Mulsant & Rey, 1871
*Oxythyrea funesta
*Protaetia (Cetonischema) speciosissima(Scopoli, 1786)
Potosia (Potosia) aeruginosa (Drury, 1773) nec Linnaeus, 1767
Subgen. Eupotosia Miksic, 1954
Protaetia (Eupotosia) affinis
(Andersch, 1797)
*Protaetia (Eupotosia) affinis affinis(Andersch, 1797)
*Protaetia (Eupotosia) affinis tyrrenicaMedvedev, 1964
Potosia (Eupotosia) affinis nat. tyrrenicaMiksic,1957
*Protaetia (Eupotosia) mirifica mirifica
(Mulsant, 1842)
Potosia (Eupotosia) koenigi balcanica Miksic, 1965
Potosia (Eupotosia) koenigi nat. balcanica Miksic, 1957
Subgen. Liocola Thomson, 1859
*Protaetia (Liocola) marmorata marmorata(Fabricius, 1792)
Liocola lugubris (Herbsts, 1786)
Subgen. Netocia A. Costa, 1852
Protaetia (Netocia) morio
(Fabricius, 1781)
*Protaetia (Netocia) morio morio(Fabricius, 1781)
Protaetia (Netocia) oblonga
(Gory & Percheron, 1833)
*Protaetia (Netocia) oblongaoblonga (Gory & Percheron, 1833)
*Protaetia (Netocia) sardea
(Gory & Percheron, 1833)
*Protaetia (Netocia) squamosa squamosa (Lefebvre, 1827)
*Protaetia (Netocia) squamosa crassicollis(Burmeister, 1842)
Subgen. Potosia
Mulsant & Rey, 1871
Protaetia (Potosia) angustata
(Scopoli, 1763)
Tropinota squalida auctt.
Aliquò, 1983
*Cetonia carthami carthamiGory & Percheron, 1833
(Germar, 1817)
12 (Fabricius, 1787)
TropinotaMulsant, 1842
AethiessaBurmeister, 1842
*Aethiessa floralis
(Kraatz, 1880)
(Poda, 1761)
(Scopoli, 1763)

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