შოთა რუსთაველის ეროვნული სამეცნიერო ფონდი biologiis 25-e saerTaSoriso olimpiadisTvis saqarTvelos nakrebi gundis wevrebis SesarCevi I turi Zvirfaso monawileebo gaqvT 2 saaTi (120wuTi) Teoriuli testis Sesasruleblad. pasuxebi CawereT pasuxebis furcelSi. pasuxi, romelic kiTxvebis furcelSi iqneba Setanili, ar Sefasdeba pasuxebi dawereT garkveviT. SesaZlebelia kiTxvas qondes erTze meti swori/araswori pasuxi da yvela ujra unda Seivsos. magaliTisTvis a b c d e maqsimaluri qula aris 40 SewyviteT pasuxebis gacema da dadeT Tqveni kalami drois amowurvisTanave. pasuxebis fiurceli da Teoriuli testebis furceli Segrovdeba weris dasrulebisas. gisurvebT warmatebebs 2 1. mitoqondrioni umetesad dakavSirebulia Semdeg funqciebTan: a. Termogenezi; b. ATP produqcia; miuTiTeT mitoqondrionis didi raodenobiT arseboba ()-niSniT, saSualo raodenobiT arseboba (-)-niSniT, xolo ar arseboba ()-niSniT. SeusabameT mitoqondrionis ZiriTadi funqcia(ebi) (a-b) Sesabamis organoebs/ujredebs. 2. arsebobs ujredebis programirebuli kvdomis e.w. “apoptozis” ramdenime meqanizmi. erT-erTi meqanizmis CarTva xdeba aqtiuri Jangbadis formiT. normaSi mitoqondrionis gareTa membranis zedapirze eqspresirdeba cila Bcl-2. sxva cilas - Apaf-1-s aqvs unari daukavSirdes Bcl-2–s. reaqtiuli Jangbadi axorcielebs Bcl-2-dan Apaf-1-is gamoTavisuflebas da, amave dros, aaqtiurebs mesame cila Bax-s, romelic, Tavis mxriv, ganWolavs mitoqondrionis membranas da gamoaTavisuflebs citoqrom C-s. gamoTavisuflebuli citoqromi C warmoqmnis kompleqss Apaf-1–Tan da caspase 9_Tan. es kompleqsi Tanamimdevrulad aaqtivebs mraval proteazas, romlebic xleCen ujredis cilebs. ras gamoiwvevs ujredSi aqtiuri Jangbadi Semdegi pirobebis SemTxvevaSi: I. ujredi eqspresirebs Apaf-1–is mutantur formas romelic yovelTvis ukavSirdeba Bcl-2–s. II. ujredebi aRar eqspresirebs Bcl-2–s. III. ujredSi xdeba Bcl-2–s overeqspresia da es cila mxolod ujredis membranisTvisaa gankuTvnili. IV. nivTiereba, romlic axangrZlivebs Bcl-2–s naxevardaSlis periods daamates ujreds. SeusabameT zemoT mocemuli mdgomareobebi (I-dan IV-mde) qvemoT CamoTvlil Sedegebs (a-dan c-mde). (2 qula) a. ujredi ewinaaRmdegeba apoptozs; b. ujredi aucileblad ganicdis apoptozs; c. ujredis bedi ucnobia. 3. G2 fazis Sesaxeb. 7.1. romeli mtkicebulebidan (ebidan) Cans, rom ujredi aris G2 fazaSi? miuTiTeT swori mtkicebuleba(ebi) ()-niSniT, xolo araswori - ()-niSniT (0,4 qula). a. himologiuri qromosomebi gaWimulia ekvatoris gaswvriv; 3 b. himologiuri qromosomebi ganizideba Sesabamisi polusebisken TiTistras ZafebiT; c. himologiuri qromosomebi jer kidev ar arian gaormagebuli; d. himologiuri qromosomebi arian haploidur anu n mdgomareobaSi. 4. qvemoT mocemulia sami saxeobis baqteriis morfologia: A (koki) Cveulebriv B (Soltiani bacila) baqteriisaTvis C (kafsuliani Cxiri) damaxasiaTebelia daukavSirdes raime zedapirs da warmoqmnas e.w. “biofilmebi”. baqteria dakavSirebis dros, sanam raime zedapirTan Seva kontaqtSi, rac ufro uaxlovdeba zdeapirs xvedeba ukugdebis zonaSi. ukugdebis zonis gadalaxavis Semdeg am sami baqteriis dakavSirebis siZliere raime zedapirTan savaraudod gansxvavebulia. SearCieT am sami baqteriisaTvis swori Tanamimdevroba dakavSirebis siZlieris mixedviT da CawereT pasuxebis furcelSi. 5. Tavdapirvelad damdgari wylis rezervuari mdidaria organuli nivTierebebiT, magram misi koncentracia male zavdeba wvimis wyliT. samive baqteria ecdeba gamonaxos saukeTeso gza gadarCes da Seeguos garemos. miuTiTeT swori mtkicebuleba(ebi) ()-niSniT, xolo araswori - ()-niSniT. a. baqteria A axdens sakvebi nivTierebebis uswrafes difuzias ujredSi. b. baqteria B SeuZlia daagrZelos Solti, rom miwvdes sakveb nivTierebebs wylis zeda fenebSi. c. baqteria C aqvs kafsula romelsac SeuZlia aqtiurad STanTqos ufro meti sakvebi nivTierebebi. 6. laborantma, mikroskopSi daTvalierebamde, sxvadasxva saRebaviT SeReba ucnobi baqteriebi. cnobilia, rom gamoyenebuli saRebavebidan (i) Rebavs lipopolisaqaridebs, (ii) birTvis garss, (iii) dnm-s, (iv) citoplazmas da (v) ribosomas. romeli saRebavi Rebavs sinjSi moTavsebul yvela tipis baqterias ganurCevlad? miuTiTeT ()-niSniT Tu SeRebavs, xolo - ()-niSniT Tu ar SeiRebavs. 4 7. SeusabameT mcenareuli struqtura (1-10) Sesabamis funqcias/niSnebs (A – J) mcenaris ujredi/qsovilis funqcia(ebi)/niSani(ebi) struqtura 1 2 Tilakoidebis membranebi vaskularuli kambiumi A wylis, momnelebeli fermentebis, sxva B centraluri vakuoli C 4 plazmodesma D 5 apikaluri meristema E 6 periderma F 7 sacriseburi mil(eb)i G 8 triqomebi H 10 ujredis meoreuli kedeli mikropile organuli da araorganuli nivTierebis dagroveba 3 9 ujredSorisi kavSirebis qseli mcenaris axali qsovilebis/organoebis warmoqmna Secvlili parenqimuli ujredebi birTvebis gareSe mcire naprali Teslkvirtze, romlis gavliTac Caizrdeba samtvre mili meqanikuri sayrdeni eleqtronebis gadamtani cilebis arseboba meoreuli gamtari qsovilebis warmoqmna I mfaravi qsovili J dacva da Sewova 8. SeiswavleT qvemoT mocemuli fesvis ganivi Wrili. 5 qvemoT mocemul cxrilSi mcenaris nawilebis aRmniSvneli ricxvebi 1-dan 18-mde SeusabameT zemoT mocemul suraTze fesvis Sesabamis nawils (A -dan H -mde) # nawili # 1 hipoderma 10 2 epiTeluri ujredebi 11 kasparis sartyeli 3 parenqimis qsilema 12 centraluri vakuoli 4 epidermuli ujredi 13 floemis parenqima 5 qsilemis boWkoebi 14 pericikli 6 fesvis bususebi 15 7 egzodermuli ujredebi nawili sklerenqimuli ujredebi Tanamgzavri ujredebi 16 floemis boWkoebi 8 qsilemis WurWlebi 17 endodermis ujredebi 9 parenqimuli ujredebi 18 kolenqimis ujredebi 9. qveviT CamoTvlilia fesvSi ionebis da wylis SeRwevis ori gza: I. simplasturi gza II. apoplasturi gza pasuxebis furcelSi mocemul suraTze aRniSneT fesvSi (garedan H –sken) ori uwyveti xaziT ionebis da wylis SeRwevis gansxvavebuli gza da TiToeul maTgans miuTiTeT (niSnebi I, II). 10. zogierTi debuleba fotosinTezis Sesaxeb mocemulia qvemoT. aRniSneT swori debuleba niSniT () , mcdari debuleba niSniT(). a. fotofosfolireba moicavs atf-s warmoqmnas/formirebas fotosinTezis sinaTlis fazaSi. b. sinaTlis mniSvnelovani roli fotisinTezis sinaTlis reaqciis dawyebisTvis mdgomareobs Tavisufali Jangbadis warmoqmnaSi. c. mcenareul ujredebSi atf sinTetazuri kompleqsi lokalizebulia mxolod Tilakoidebis membranebSi. d. dReisaTvis Tvlian rom, CO2-is SaqarSi damafiqsirebeli fermentebi lokalizebulia qloroplastis stromaSi. 6 11. SeusabameT aRwerilobasa an Sedegs (A – J) maTi Sesabamisi terminebi (1 – 10). termini aRwera/Sedegi organizmis fiziologiuri reaqcia dRis an Ramis 1 eTileni A 2 fotoperiodizmi B gverdiTi kvirtebis zrdis SeCereba C Reros dagrZeleba dauzianebel mcenareebSi 3 apikaluri dominireba xangrZlivobaze. dabali temperaturis xangrZlivi moqmedeba xels 4 Tigmotropizmi D 5 filotaqsia E 6 citokinini F 7 giberelini G foTlebis ganlageba Reroze 8 statoliTi H mcenaris sapasuxo reaqcia Sexebze 9 vernalizacia I daberebis procesis SeCereba 10 heliotropizmi J graviotropizmi uwyobs mcenaris ayvavebas iwvevs foTlebis da nayofebis cvenas mzardi Reroebis mzisken an sinaTlis sxva wyarosken gadaxra 12. qvemoT mocemulia oTxi sxvadasxva cxovelis A – D sunTqvis sixSire, guliscemis sixSiris da sxeulis temperaturis monacemebi. sunTqvis sixSire guliscemis sixSire sxeulis temperatura (CasunTqva/wT) (guliscema /wT) (°C) A 160 500 36.5 B 15 40 37.2 C 28 190 38.2 D 8 28 35.9 cxoveli daalageT cxovelebi (A-D) zedapiris farTobis sxeulis moculobasTan fardobis klebadobis mixedviT. 13. daalageT cxovelebi (A-D) (me-12 kiTxvidan) sxeulSi sisxlis mTliani moculobis zrdadobis mixedviT. 7 14. sxvadsxva tipis cxovelis gazTa cvlaSi CarTulia sunTqvis sxvadsxva organo (a – d) da sisxlis mimoqcevis sistema (Ria da daxuruli). TiToeuli cxovelisTvis aRniSneT sisxlis mimoqcevis sistemis tipi. (niSniT () Ria sistema da niSniT (X) sisxlis mimoqcevis daxuruli sistema) da aseve SeusabameT Sesabamisi organoebi (a – d) CamoTvlil zrdasrul cxovelebs. a. filtvebi b. layuCebi c. kani d. traqea 15. ZuZumwovrebis mier gamoyofili nerwyvis raodenoba damokidebulia Wamisas ReWvis intensiurobaze. SeusabameT cxovelebi (a – d) pasuxebis furcelSi moyvanil cxrilSi mocemul gamoyofili nerwyvis raodenobas a. mgeli b. cxeni c. msxvilfexa rqosani pirutyvi d. adamiani 16. alergia aris hipermgrZnobire adamianebis imunuri sistemis reaqcia, romelic miiRweva organizmSi antigenis ganmeorebiTi SeRwevis dros. fsevdoalergia gavs alergias klinikuri niSnebiT, magram gansxvavebaa, is rom mis ganviTarebaSi imunuri sistema ar monawileobs. mocemulia paTologiuri procesebi: a. IgE klasis antisxeulebis saerTo raodenoba izrdeba plazmaSi. b. plazmaSi aRiniSneba IgE klasis specifikuri antisxeulebi. c. gamoTavisufldeba anTebis ZiriTadi mediatori - histamini. d. reaqciis gamosavlenad antigenebis minimaluri raodenobaa sakmarisi. aRniSneT romeli procesia(ebia) marTebuli alergiisTvis da romeli fsevdoalergiisTvis niSniT () xolo ara marTebuli mtkicebulebebi aRniSneT niSniT (X) pasuxebis furcelSi . 17. evropuli (mtknari wylis) gvelTevza Jangbads Cveulebriv layuCebiT Rebulobs, maram SeuZlia wylis gareSe yofna kaniT sunTqvis unaris gamo. mocemuli grafiki uCvenebs JangbadiT sisxlis gajerebas da sxvadsxva organoebisTvis Jangbadis miwodebas, roca gvelTevza ar aris wyalSi. (pirobiT erTeulebSi): 8 O2 done (pirobiTi erTeuli) dro (pirobiTi erTeuli) A, ▲B, ■ C, ●D SeusabameT CamoTvlili mtkicebulebebi (I - IV) Sesabamis mrudebs (A – D) I. sisxlis totaluri gajereba JangbadiT. II. Jangbadis miwodeba layuCebiT. I. Jangbadis miwodeba kaniT. II. Jangbadis miwodeba sacuravi buStiT 18. cxovelebis anatomiuri maxasiaTeblebi adaptirebulia kvebis sxvadasxva tipTan (a – d). a. xorcismWamelebi b. nairmWamelebi c. aramcoxneli balaxismWamelebi d. mcoxneli balaxismWamelebi SeusabameT kvebis sxvadsxva tipebi (a – d) kbilebis Sesabamis maxasiaTeblebs (I – IV). I. zeda mWrelebis ararseboba, kbilis firfitis arseboba, ZiriTadi kbilebi moZraobs mxolod erT sibrtyeSi horizontalurad. II. eSvebi kargad aris ganviTarebili da gamoiyeneba sakvebis dasaglejad. III. ZiriTad kbilebi saReWi kbilebia. 9 IV. mWrelebi gankuTvnilia sakvebis mosaWideblad, ZiriTadi kbilebi ganlagebulia garkveuli kuTxiT, ybebi moZraobs cirkularulad (vertikulurad da horizontalurad) 19. saWmlis momnelebli sxeulis zedapirTan sistemis (sms) zedapiris farTobis Sefardeba mTliani balaximWamel, gansxvavebulia. SeusabameT arsebul cxrilSi moyvanil nairmWamel da xorcismWamelebs kvebis sxvadsxva tipebi (a – d) saWmlis momnelebli sistemis Soris pasuxebis furcelSi zedapiris fardobas mTliani sxeulis zedapirTan 20. saWmlis momnelebeli sistemis adaptacia (a – c) SeusabameT Sesabamis anatomiur aRwerilobas . anatomiuri aRweriloba: I. martivi kuWi, silosis utilizaciis SezRuduli unari II. martivi kuWi, silosis utilizaciis uunaroba III. kargad ganviTarebiuli tomriseburi kuWi, silosis yovelmxrivi da efeqturi utilizaciis unariT kvebiTi adaptacia: a. pirveladi monelebis Semdeg duRili da Sewova. b. duRili pirvelad monelebamde da Sewovamde. c. marcvlebis, xilis da bostneulis zogierTi nivTierebis monelebis SeuZlebloba. 21. xerxemlianebis yvela klasSi gvxvdeba ramdenime saxeoba, romelTac xandaxan mainc SeuZliaT haerSi asvla (mag. “mfrinavi” Tevzi, bayayi, xvliki da ciyvi). maT ar SeuZliaT namdvili frena, magram daRmasvlis Sesaneleblad iyeneben iseT unarebs rogoricaa livlivi da paraSutismagvari moZraoba. molivlive cxovelebi amcireben muxruWs (D) minimumamde da iyeneben amwev Zalas (L) rom warmoqmnan xelsayreli damokidebuleba amwev Zalasa da muxruWs Soris (fardoba L/D). amis sawinaaRmdegod, cxovelebi, romlebic paraSutireben, zrdian D-s, radgan maT umravlesad aqvT mniSvnelovnad naklebi zedapiris farTobi imisTvis rom warmoqmnan L. roca cxovelebi miaRweven stabilur livlivs, masze moqmedebs ramdenime Zala. (ix. suraTi qvemoT). gaSlili sxeulis winaaRmdegoba (R) haeris nakadTan warmoqmnis L-s. amasTan, masze moqmedebs damuxruWeba (D) moZraobis sawinaaRmdego mimarTulebiT da, aseve, sxeulis wona (mg). daRmasvlis traqtoria warmoqmnis kuTxes () miwasTan. 10 pasuxebis furcelSi cxrilSi motanili savaraudo L/D da sidideebis gaswvriv miuTiTeT Sesabamisi cxoveli (a an b). 22. mfrinav xvlikebSi patagilumi warmoadgens kanis nakecs, romelic damagrebulia wina da ukanakidurebs Soris. Russell and Dijkstra (2001) Seadares patagiumi da damatebiTi aerodinamikuri danamatebi xvlikebis or sxvadasxva saxeobaSi Draco volans (mfrinavi drakoni) da Ptychozoon kuhli (mfrinavi gekoni) ------ Draco volans; Ptychozoon kuhli 11 aRniSneT niSani ()-T swori daskvna(ebi) warmodegnili xvlikebis haerSi gadasaadgileblad arsebuli morfologiuri adaptaciis Sesaxeb. xolo araswori daskvnebi ki niSniT (X). a. miuxedavad imisa, rom D. volans saSualo wona ufro mcirea, vidre P. kuhli-s, maTi sxeulis farTobis Sefardeba wonis erTeulTan erTnairia. b. potagilusis farTobis Sefardeba masasTan gviCvenebs, rom P. kuhli potagilusi gacilebiT didia, vidre eqvivalenturi masis D. volans potagilusi. c. damxmare struqturebi ufro mniSvnelovan gavlenas axdenen D. volans-is saerTo aerodinamikur farTobze vidre P. kuhli-s. d. am ori saxeobis mTliani sxeulis farTobi TiTqmis erTnairia. e. proporciuli farTobi, romelic ganpirobebulia potagilumiT ufro didia D. volans-Si, vidre P. kuhli-Si, romelic kompensirdeba damatebiTad kudis da nakecebis arsebobiT. 23. hipofizSi ramdenime maregulirebeli peptidi sinTezirdeba propeptididan - proopiomelanokortinidan (POMC). POMC ixliCeba proteolizurad (A – E) sxvadsxva produqtebad. qvemoT yvela peptidi warmodgenilia N boloTi marcxniv da C boloTi marjvniv. SeniSvna: TiToeuli fermenti xleCs mxolod “momwifebul” prekursors. dawereT minimum ramdeni fermentia saWiro, rom POMC-dan miRebuli iqnas β –MSH. 24. dawereT minimum ramdeni fermentia saWiro, rom POMC-dan miRebuli iqnas α -MSH. 25. mamri qelipedebs) kiborCxala signalebis iyenebs Tavis gadasacemad gadidebul (mag. brZola qelipedebs/marwuxebsM(mTavar teritoriisTvis, mdedris yuradRebis mipyroba da sxv.). studenti swavlobda urTierTobebs mdedr da mamr kiborCxalebs Soris. studenti Seswavlis dros iyenebda sarkeebs, romlebic mamri 12 kiborCxalas ori sxvadasxva zomis gamosaxulebas iZleoda. eqsperimentSi gamoyenebuli sarkeebis kombinacia Semdegnairi iyo: 10x : 3x (I cda), 3x : 1x (II cda) da 10x : 1x (III cda). TiToeuli cdis msvlelobisas 20 mdedrs uCvenebdnen 10 mamrs (TiToeul cdas imeorebdnen 3-jer). studenti iwerda/afiqsirebda Sedegebs: mdedrebis procents (grafiki A) da dros, romelic sWirdeboda mdedrs yvela gamosaxulebamde misaRwevad (grafiki B) TiToeul cdaSi, agreTve, mamri iyenebda marcxena marwuxs % dro (wm) (memarcxene iyo) Tu marjvenas (memarjvene) (grafiki C). cda % cda cda metad gadidebuli gamosaxuleba memarjvene mamri mcired gadidebuli gamosaxuleba memarcxene mamri miuTiTeT swori daskvna (ebi), romlis gamotana aris SesaZlebeli mamri da mdedri kiborCxalebis damokidebulebis Sesaxeb da aRniSneT niSniT (), xolo araswori pasuxebi aRniSneT (x)-iT. a. mocemuli saxeobis mdedrebi upiratesobas aniWeben didi zomis mamrebs. 13 b. SejvarebisTvis arCevanSi mamris memarjveneoba/memarcxeneoba aris ZiriTadi kriteriumi. c. mamrebi, romlebic ufro swrafad amoZraveben marwuxebs, rogorc wesi meti mdedris yuradRebas iqceven. d. saSualo dro, romelic sWirdeba mdedrebs arCevanis gasakeTeblad gansxvavdeboda II da III cdebSi.. e. SesamCnevi sxvaoba mamrebis marwuxebis zomaSi SesaZloa iyos aucilebeli piroba mdedrisTvis gadawyvetilebis misaRebad. 26. wylis frinveli didi Cvama (Phalacrocorax carbo) ikvebeba TevziT. isini wyalSi yvinTaven, da mxedvelobis gamoyenebiT pouloben msxverpls, amitom didi mniSvneloba eniWeba wylis sisufTaves. ufro xSirad Cvama individualurad nadirobs, magram wylis arasakmarisi gamWvirvaleobis dros maT SeuZliaT gamoimuSaon meTodebi erToblivi jgufuri nadirobisas. (1.2 qula). 1-1,5m sxva Cvamebi Wamen Tevzs mRvrie wyali 4m gamWvievale wyali Cvamebi morekaven Tevzs aRniSneT niSniT () process(eb)i, romlebsac aqvT mniSvneloba erToblivi nadirobis strategiis SemuSavebisTvis da niSniT() proces(eb)i, romlebsac msgavsi mniSvneloba ar aqvT. a. konkurencia; b. miCveva; c. jgufuri/socialuri daswavla; d. imprintingi; e. mcdeloba da Secdomebi. 14 27. moaxdines 28 wyvili avtosomuri qromosomis da wyvil sasqeso qromosomis (XY) Semcveli atlantikuri mikroineqciis teqnikis oragulis (Salmo gamoyenebiT salar) genomis warmatebulad sekvenireba. moxda zrdis dnm-s hormonis transgenis CaSeneba oragulis embriolnis zigotis stadiaze. Sedegad, miRebuli iqna 4 transgenuli idividi (F0 mSobeli) 2 mamri da 2 mdedri. zrdis hormonis transgeni regulirdeba RviZlis specifikuri enhanseriT da oTxive transgenul mSobels aqvs zrdis hormonis maRali done plazmaSi, rac xels uwyobs daCqarebul zrdas. dadasturebulia, rom am individebSi transgenis erTi aslis CaSeneba (insercia) xdeba. daCqarebuli zrdis mqone transgenuli oragulebis mdgradi xazi gamoyvanili iqna SejvarebiT. pirveli Taobis (F1) misaRebad, orive mamrobiTi da mdedrobiTi transgenuli individi (F0) Seajvares Sesabamisi sqesis velur, aratransgenul oragulTan. F2 Taobis dasadgenad Tqven unda moaxdinoT axlonaTesauri Sejvareba (da-Zmas Soris), raTa gamoavlinoT homozigoturi transgenuli individebi, romlebic atareben zrdis hormonis transgens. rogori iqneba %-Si mosalodneli genotipis Tanafardoba. 28. qaTami, mokle frTebiT da fexebiT (kreperi), Seajvares normalur qaTams. Semdgom TaobaSi miiRes erTnairi raodenobis kreperi da normaluri qaTami. rodesac Seajvares kreperebi erTmaneTs, miiRes 2 kreperi da 1 normaluri qaTami. normaluri qaTmebis SejvarebiT miiReba mxolod normaluri Taoba. ra aris umartivesi genetikuri safuZveli kreperebisa da normaluri qaTmebisaTvis? miuTiTeT swori pasuxi(ebi) ()-niSniT, xolo araswori niSani(ebi) - ()-niSniT. 29. zRvis goWis Savi bewvi ganpirobebulia B dominanturi geniT, xolo TeTri misi recesiuli aleliT b. vTqvaT, II 1 da II 4 ar arian recesiuli alelis matareblebi ra aris albaToba imisa, rom II 3 aris heterozigoti? 15 30. ra aris albaToba imisa, rom III 1 x III 2 Sejvarebidan erTi mainc iqneba TeTrbewviani? 31. erT-erT marcvlovan mcenareTa kulturaSi, ramdenime geni marTavs anTocianinis gamomuSavebas. anTocianinis ararsebobis dros aRmonacens aqvs mwvane foTlebi, anTocianinis arsebobisas, ki foTlebs gadahkravT iasamnisferi. genis lokusi, romelsac uwodeben C1, moqmedebs rogorc transaqtivirebuli induqtoruli lokusi, sul mcire ori lokusisTvis (ChsA and ChsJ), romelic anToceaninis sinTezSi monawile or fermentebs kodireben. anToceaninis sinTezi homozigotur recesiul genotipSi blokirebulia, rogorc naCvebnebia suraTze chsA-Tvis (qvemoT). DdavuSvaT, rom arsebobs Semdegi mdgomareobebi: (i) samive lokusisaTvis misaRebia damoukidebeli ganawileba da (ii) 3 lokusidan nebismieris recesiuli aleliT homozigoturoba ganapirobebs aRmonacenis mwvane fers. miuTiTeT mosalodneli fenotipuri STamomavlobis swori Tanafardoba ori SejvarebisTvis cxrilSi pasuxebis furcelSi ()-niSniT, xolo Seusabamo ki - ()-niSniT. Chs A da Chs J induqtori Sualeduri A iasamnisferi pigmenti Sualeduri B fermenti ChsJ ChsA fermentis ararseboba 32. CanTosani Txunela (rigi Notoryctemorphia) binadrobs samxreT avstraliis udabnoSi, dasavleT avstraliasa da CrdiloeT teritoriaze. isini miwaSi gvirabebis gaTxris dros qviSas ukan yrian, TiTqos miwaSi micuraven. Tvalebi mTlianad dafarulia kaniT, 1 mm diametriT da ar gaaCnia broli an guga. mxedvelobis nervi Zlier reducirebulia. miuTiTeT swori mtkicebuleba(ebi) (a-dan – c-mde) ()- niSniT, xolo araswori ki - ()-niSniT. a. brolis ar arseboba aris homologia gamoqvabulis buzis TvalSi omatidias (mozaikuri Tvalis patara nawili) ararsebobisa. b. Zlier reduqcirebuli optikuri nervi aris rudimenturi srtuqtura. c. Txunelas Tvalebi aris kengurus Tvalebis analogi. 16 33. interfotoreceptoris retinoidis damakavSirebeli cila (IRBP) aris genis erTi asliT warmodgenili. ZuZumwovrebSi misi produqti mniSvnelovan rols asrulebs rodopsinis aRdgenaSi. am genis dnm-s Tanamimdevrobebi Sedarebuli iqna ramdenime TxunelaSi. IRBP makodirebeli genis aratranskribirebadi jaWvi naCvenebia qvemoT. gaiTvaliswineT, rom genis dasawyisi ar aris naCvenebi da naCvenebia genis CamokiTxvis swori CarCo. im kodonidan dawyebuli, romelic monawileobs CarCos gadawevis mutaciaSi, dawereT Tanamimdevrulad aminomJavebi, romelsac es geni kodirebs vombatebSi da CanTosan TxunelebSi. gamoiyeneT genetikuri kodis cxrili. 34. miuTiTeT swori mtkicebuleba ()-niSniT, araswori ki - ()-niSniT. vombatebisgan gansxvavebiT, CanTosan TxunelebSi IRBP genis produqti: a. Seicavs mraval Secvlil aminomJavas. b. aris damoklebuli, radgan Seicavs Stop kodons adreul wertilSi. 17 35. 1980 wels moulodnelad amoifrqva mTa st. elena vaSingtonis StatSi. amofrqvevis Semdeg darCa landSafti sakvebi nivTierebebiT Raribi, gaZlierebuli gvalviT da xSirad moZravi gruntiT. amofrqvevis Semdeg Tanasazogadoebis aRdgenis Seswavlis mizniT krateris garSemo xeebis gavrcelebis zeda zRarze sxvadsaxva adgilas daadgines sacdeli nakveTebi. suraTi gviCvenebs saxeobebis raodenobas da safaris procentobas miRebuls erT-erTi nakveTidan 1981wlidan 1998 wlamde. % safari saxeobis raodenoba saxeoba safari wlebi aRniSneT niSniT () swori daskvnebi, romlebic SeiZleba am suraTidan gamomdinareobdes, mcdari daskvnebi aRniSneT (X) a. amofrqvevam gaanadgura mcenareuli safari xeebis gavrcelebis zeda zRvris zemoT. b. arc sivrce da arc sinaTle ar aris SemzRudveli faqtori am areSi. c. mxolod ramdenime damatebiTi saxeobebi gamoCndnen 1982 wlidan. d. mTliani mcenareuli safari am areSi gaizarda SedarebiT nela vulkanuri danaleqebis mkacr pirobebSi. e. am kvlevaSi miaRwies iqna stabiluri 20 saxeobisagan Semdgari mcenreuli Tanasazogadoebis Camoyalibebas, rac sukcesiis procesSi klimaqteruli etapis dadgomas miuTiTebs. 18 36. sqema qvemoT gviCvenebs zRvis Tanasazogadoebis simulacias. sazogadoebaSi populaciebs Soris urTierTobis Seswavlis mizniT. normaluri mdgomareoba zRvis varskvlavi mocilebulia Sedegi Sua litorali qveda litorali Starfish - zRvis varskvlavi; mussels - midiebi – orsagdulianebi; Barnacles – fexsaxsriani; Gooseneck barnacles – fexsaxsriani; Rockweed – mura wyalmcenareebi am suraTis mixedviT aRniSneT niSniT () swori daskvnebi, mcdari daskvnebi aRniSneT (X) a. sazogadoeba bunebriv mdgomareobaSi Seicavs 4 saxeobas cxovelebis samefodan. b. suraTze warmodgenilia mxolod: kaneklianebi, moluskebi da fexsaxsrianebi. c. TavianT bunebriv garemoSi zRvis varskvlavi ZiriTad saxeobas ar warmoadgens. d. TavianT bunebriv garemoSi midiebis simkvrive didia Sua litoralur zonaSi da mcirea qveda litoralurSi, radgan zRvis varskvlavi cxovrobs qveda litoralur zonaSi. e. kvlevis bolo etapze sazogadoeba qreba da mxolod erTi saxeoba zrdis Tavis ekolgiur niSas. f. demonstrirebulia sxva populaciis konkurentuli gacxrilva midiebis mier. g. midiebma daipyres fundamenturi niSa orive litoraluli zonis (Sua da qveda) CaTvliT. h. bunebrivi mdgomareba moicavs biotur urTierTobas rogoricaa saxeobaTaSorisi konkurencia da mtacebloba. 19 37. kladistikuri sistematikuri mkvlevarebi iyenebs sxvadsxva jgufebis Sedarebas raTa moaxdinos warmoebuli niSnebis da jgufebis saerTo sawyisi niSnebis diferencireba. amisTvis isini iyeneben sxva jgufs, romlic axlos dgas maT mier Sesaswavl jgufTan. piroba: sxva jgufi unda iyos nakleb naTesauri sacdeli jgufis wvrebTan vidre sacdeli jgufis wevrebi erTmaneTTan. hipoTeza: sawyisi niSnebi, romelic win uswrebs jgufebis divergencias - homologiuria. am Teoriuli koncepciis mxedvelobaSi miRebiT, zogierTma mecnierma Seiswavla sxva jgufi. monacemebi mocemulia cxrilSi: cxovelTa jgufebi niSnebi A1 A2 A3 A4 A5 A6 a. Tma 0 0 0 0 1 0 0 1 0 0 1 0 c. moZraobisTvis 4 fexi 0 1 0 0 1 1 d. SeerTebuli ybebi 0 1 1 0 1 1 e. xerxemali 0 1 1 1 1 1 f. 1 1 1 1 1 1 b. amnioturi kvercxi damatebiTi Canasaxovani membranebiT notoqordi SeniSvna 0: niSnis ararseboba, 1: niSnis arseboba zemoT moyvanili informaciis analizis Semdeg AdaadgineT (A1-A6), romelia “sxva” jgufi. 38. miuTiTeT niSnebi (a-f), romelic saerToa sxva da sacdeli jgufisTvis. 39. miuTiTeT sawyisi niSani, romelic sacdeli jgufis yvela wevrs aqvs. 40. aRniSneT saboloo divergenciis ganmsazRvreli niSani. 20
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